Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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Catalogue of Palaearctic Coleoptera - Volume 1, 2003

I. Lobl & A. Smetana (Apollo Books)

Index to species‑group names
- the order is by species-group name and then by current genus rather than author

- species-level synonyms are in Italics

- the original combination is in square brackets following the current generic combination

-junior homonyms which were not indicated as such in Volume 1 are marked [HN], or [HN!] if currently valid, but not yet dealt with

-names which were cited incorrectly in Volume 1 are in (parentheses), preceded by the corrected spelling
aabadensis Lassalle, 1991, Carabus 193

aanistschenkoi O. Berlov & E. Berlov, 1999, Pterostichus 486

aapsorum Belousov, 1991, Pterostichus 500

abaciformis Straneo, 1955, Pterostichus 497

abacoides Lucas, 1846, Orthomus [Oodes] 476

abadiei A. Gaudin, 1947, Hydraphaenops 311

abadiei A. Gaudin, 1947, Pterostichus [Lianoe] 498

abadzechus Zamotajlov, 1988, Carabus 201

abagonensis Stark, 1894, Carabus 200

abagonensis Reitter, 1896, Pterostichus 495

abakumovi Belousov & Kabak, 1996, Trechus 325



abalisensis Morvan, 1977, Ophonus [Metophonus] 391

abantayaginiensis Schweiger, 1962, Carabus 198

abanticola Korge, 1964, Poecilus [Pterostichus] 482

abasarukini O. Berlov & E. Berlov, 1996, Pterostichus 487

abashicus Belousov, 1998, Troglocimmerites 344



abasinus Rost, 1893, Carabus [Plectes] 200

abasinus Rost, 1892, Harpalus 374

abasinus Belousov, 1991, Pterostichus [HN, repl. in vol. 2] 504

abasinus Rost, 1892, Pterostichus [Aphaonus] 486

abaxoides Brullé, 1839, Calathus 526

abaxoides Dejean, 1828, Pterostichus [Feronia] 489

abbasi Ali, 1978, Cephalota [Cicindela] 102

abbreviata Chaudoir, 1842, Amara [Celia] 555

abbreviatum Solsky, 1874, Bembidion 260

abbreviatus Fabricius, 1787, Agabus [Dytiscus] 37

abbreviatus Brullé, 1835, Carabus 176



abbreviatus Chaudoir, 1846, Dyschirius 228

abbreviatus Wehncke, 1880, Haliplus 31



abbreviatus Motschulsky, 1850, Molops 472

abbreviatus Dejean, 1825, Scarites 232

abchasicum Müller‑Motzfeld, 1989, Bembidion 245

abchasicus Motschulsky, 1850, Carabus 187


abchasicus Reitter, 1896, Carabus 200

abchasicus Reitter, 1896, Pterostichus 504

abcuma Uéno, 1992, Trechiama 321

abdelkrimi Netolitzky, 1926, Bembidion 264

abditus Fedorenko, 1993, Dyschirius [Dyschiriodes] 228

abditus Lutshnik, 1934, Poecilus [Platysma] 480

abdominalis A. Costa, 1847, Agabus [Colymbetes] 38

abdominalis Ragusa, 1887, Agabus 40

abdominalis Motschulsky, 1844, Amara [Bradytus] 551

abdominalis Motschulsky, 1844, Dicheirotrichus [Ophonus] 402

abdominalis Reiche & Saulcy, 1855, Leistus 84

abdominalis Gené, 1836, Stenolophus 405

abdominalis Motschulsky, 1844, Trechoblemus [Trechus] 324

abdurakhmanovi Zamotajlov, 1994, Deltomerus 282

abdurakhmanovi Belousov, 1990, Trechus 325

abei Nakane, 1997, Panagaeus 447

abei Morita, 1997, Trichotichnus 395

abeillei Bedel, 1879, Bembidion 251

abeillei Jeannel, 1942, Cymindis 411

abeillei Brisout de Barneville, 1885, Microlestes [Blechrus] 422

abeillei Pandellé, 1872, Trechus 325

aberanus Jedlika, 1955, Platynus [Colpodes] 464

aberrans Baudi di Selve, 1864, Amara 555

aberrans Uéno, 1978, Kurasawatrechus 313



aberrans A. Morawitz, 1862, Pterostichus 487

aberrans Tschitschérine, 1906, Trichotichnus [Amaroschesis] 394

aberratus Gemminger & Harold, 1868, Rhantus 46

abishirensis Belousov, 1991, Pterostichus 504

abkhazicus Belousov, 1998, Nannotrechus 314

abkhazicus Jeannel, 1960, Trechus 330

abnormale Jedlika, 1960, Agonum 454

abnormale Jedlika, 1965, Bembidion 254

abnormalis Jedlika, 1956, Amara 548

abnormalis Jedlika, 1958, Laemostenus 539

abnormalis Jedlika, 1968, Nebria 86

abnormalis Morita, 1998, Nipponosynuchus 544

abnormalis Jedlika, 1961, Pseudotaphoxenus [Taphoxenus] 540

abnormicolle Puel, 1938, Agonum 451

abnormicolle Heyden, 1882, Asaphidion [Tachypus] 242

abnormicollis Ballion, 1871, Agabus 38

abnormicollis Semenov, 1900, Deronectes 57

abnormis Tschitschérine, 1894, Amara 561

abnormis J. R. Sahlberg, 1900, Corcyranillus [Anillus] 238

abnormis Knirsch, 1913, Duvalius [Anophthalmus] 298

abnormis J. R. Sahlberg, 1880, Pterostichus [Feronia] 505



abodiensis Dupré, 1988, Aphoenops 293

aborana Andrewes, 1925, Nebria 90

abreshensis Lutshnik, 1916, Tapinopterus [Platysma] 519

abrubtesculta W. Horn, 1925, Tricondyla 118



abruzzensis Schauberger, 1923, Amara 559

absentivus Belousov & Kabak, 1993, Carabus 182

absoloni Obenberger, 1917, Nebria 86

abstersus Bates, 1873, Chlaenius 354

abudarini O. Berlov & E. Berlov, 1997, Pterostichus 493

abyssinica Chaudoir, 1876, Tachyura [Tachys] 278

academiae Deuve, 1992, Trechus 325

acaroides Waltl, 1838, Sphaerius 26

accentifer Raffray, 1873, Mesolestes [Dromius] 421

accentifera Zoubkoff, 1833, Cymindis 411

accessor Wrase, 1997, Psychristus 404

accipitris Uéno, 1983, Trechiama 321

acco Uéno, 1989, Trechiama 321

acco Uéno, 1991, Trechus 325

accuratum Casey, 1924, Bembidion 254

accuratus Chaudoir, 1848, Carabus 198

acheicus Zamotajlov, 1991, Carabus 132

acherontius Schaufuss, 1881, Duvalius [Anophthalmus] 307

achillecasalei Deuve, 1997, Carabus 189

achillecasalei Deuve, 1998, Trechus 325

achilles G. Hauser, 1924, Carabus [Apotomopterus] 124

achilles Sciaky, 1987, Ophonus 389

achunensis Gottwald, 1985, Carabus 132

aciculatus Herbst, 1784, Cybister [Dytiscus] 50

aciculatus Schaum, 1864, Zabrus [Eutroctes] 568



acolpomatus Antoine, 1941, Harpalus 375

acorep Lassalle & Prunier, 1993, Carabus 125

acosmos Cavazzuti, 1997, Carabus 125

acragas De Monte, 1956, Bembidion 20, 249



acrocolius Beck, 1817, Ocys [Bembidion] 273

acroglyptus Bates, 1892, Platynus [Colpodes] 464

acrogonus Chaudoir, 1843, Pterostichus [Orthomus] 489

actenipoides Lassalle, 1993, Laemostenus 533



acuductus Marsham, 1802, Agabus [Dytiscus] 38

aculeatus Jeanne, 1976, Leistus 81

aculeatus Reiche & Saulcy, 1855, Pachycarus 367



acuminata Casey, 1918, Amara 549

acuminata Paykull, 1798, Amara [Carabus] 549

acuminata Kollar, 1836, Cylindera [Cicindela] 111

acuminatellus Fairmaire, 1876, Nebrioporus [Hydroporus] 67

acuminatus Wollaston, 1862, Calathidius [Calathus] 532


acuminatus Ménétriés, 1836, Carabus 182

acuminatus Motschulsky, 1844, Harpalus 380

acuminatus Stephens, 1835, Harpalus 378

acuminatus Sturm, 1835, Hydroporus 60

acuminatus Motschulsky, 1859, Hydrovatus 70

acupalpoides Reitter, 1900, Harpalus 371

acupalpoides Bates, 1883, Perigona 448

acuspinus Tschitschérine, 1901, Pterostichus [Platysma] 507

acutangula Chaudoir, 1873, Cymindis 412

acutangulus Chaudoir, 1876, Aptinus 212

acutangulus Chaudoir, 1879, Carabus 197



acutangulus Eichler, 1924, Carabus 199

acutangulus C. G. Thomson, 1856, Hydroporus 60

acutangulus Schaufuss, 1862, Laemostenus [Pristonychus] 532

acutangulus Perrault, 1979, Leistus 81

acutangulus Wollaston, 1857, Olisthopus 461

acutangulus Andrewes, 1923, Onycholabis 461



acutangulus Jeannel, 1947, Pterostichus [Haptoderus] 492

acutangulus Semenov, 1908, Taphoxenus 544

acutangulus Apfelbeck, 1902, Trechus 326

acutangulus Mateu, 1963, Trymosternus 410



acutesculptus Chaudoir, 1877, Carabus 174

acuticauda Casey, 1924, Amara 549

acuticolle Motschulsky, 1860, Bembidion [Peryphus] 265

acuticollis Putzeys, 1873, Calathus 524

acuticollis Fairmaire, 1889, Callistomimus [Callistus] 348

acuticollis Motschulsky, 1858, Carabus 188



acuticollis L. Dufour, 1857, Cychrus 204

acuticollis Putzeys, 1866, Notiophilus 97

acuticollis L. Dufour, 1843, Perileptus [Blemus] 288

acuticollis Sciaky & Pavesi, 1994, Trechus 325

acutidens Westwood, 1833, Platyrhopalus 211

acutidens Fairmaire, 1889, Pterostichus [Omaseus] 495

acutidens Chaudoir, 1855, Scarites 232

acutidens Uéno, 1986, Trechiama 321



acutipalpis Putzeys, 1877, Syleter [Psilus] 221

acutipennis Küster, 1847, Parophonus [Harpalus] 392

acutipennis Bates, 1892, Progonochaetus [Gnathaphanus] 363

acutithorax Deuve, 1989, Carabus 152

acutithorax Casale, 1988, Pseudotaphoxenus 540

acutum Marsham, 1802, Bembidion [Carabus] 251

acutus Lapouge, 1904, Carabus 193

acutus Reitter, 1893, Daptus 369



acutus Putzeys, 1866, Dyschirius 225

acutus Sharp, 1882, Hydrovatus 70

adachii Kamiya, 1932, Hydrovatus 71

adamantina Kolenati, 1845, Amara 550

adamellensis Jeannel, 1926, Duvalius 306

adamellicola Ganglbauer, 1904, Carabus 180

adamsi M. F. Adams, 1817, Carabus 192



adamsianus Mandl, 1975, Carabus 192

adamsii Fischer von Waldheim, 1817, Carabus 201

adamsii Clark, 1864, Graphoderus [Hydaticus] 49



adamsii Fischer von Waldheim, 1817, Zabrus 568

adanae De Monte, 1953, Bembidion 253

adanensis Jedlika, 1961, Apotomus 237

adanensis Maan, 1941, Leistus 84

adanensis Schauberger, 1933, Ophonus [Harpalus] 391

adangensis Gottwald, 1983, Carabus 137

adatarasanus Ishikawa, 1966, Carabus [Leptocarabus] 149

adductus Cavazzuti & Ratti, 1998, Carabus 124

adehinotum Lassalle, 1992, Calosoma [Callisthenes] 118

adelaidae Stark, 1889, Carabus [Neoplectes] 133

adelaidae Reitter, 1899, Pterostichus 500

adelioides W. S. MacLeay, 1825, Aephnidius [Anaulacus] 357

adelosia Andrewes, 1934, Synuchus [Calathus] 545

adelphus Rost, 1892, Carabus [Plectes] 200



adelus Iablokoff‑Khnzorian, 1973, Brachinus 215

ademidovae O. Berlov, 1996, Pterostichus 487

adenticulatus Huang, 1992, Harpalus 384

adjamdehensis Morvan, 1974, Trechus 339

adjarica Shilenkov, 1983, Nebria 84



adjunctus Csiki, 1927, Carabus 124

adolescens G. Hauser, 1925, Carabus [Apotomopterus] 123



adonis Hampe, 1853, Carabus 138

adonis Tschitschérine, 1895, Platynus [Colpodes] 464



adoxus Fischer von Waldheim, 1823, Carabus 198

adoxus Andrewes, 1923, Philorhizus [Dromius] 425

adpressus Aubé, 1837, Agabus 38

adriaenssensi Lassalle & Prunier, 1993, Carabus 143

adriaenssensi Lassalle, 1998, Omphreus 446

adriatica J. Müller, 1921, Cymindis 411



adspersus Gmelin, 1790, Carabus 160

adspersus Panzer, 1796, Rhantus [Dytiscus] 46

adstrictus Eschscholtz, 1823, Pterostichus 487

adsypschi Gottwald, 1985, Carabus 132

adunca Gravenhorst, 1807, Cylindera [Cicindela] 110

adusta L. Redtenbacher, 1843, Cymindis 411



adusticauda G. Costa, 1907, Bembidion 249

adustipennis Reitter, 1892, Apotomus 237

adustum Schaum, 1860, Bembidion 254

adustus Guignot, 1954, Agabus 38

adustus Jeannel, 1962, Trechus 325

advena Schauberger, 1921, Brachinus 213

advena Wollaston, 1862, Calathus 526

advena Quensel, 1806, Poecilus [Carabus] 480

advena Uéno, 1983, Trechiama 321

adventicius Schauberger, 1934, Harpalus 380

adyg Zamotajlov, 1997, Carabus 137

adygorum Belousov & Sokolov, 1996, Bembidion 258

aeacus Saulcy, 1864, Aphoenops [Anophthalmus] 294

aeacus L. Miller, 1861, Laemostenus [Sphodrus] 534

aedeagalis Straneo, 1989, Pterostichus 509

aegaeus Apfelbeck, 1904, Zabrus 570

aeger Andrewes, 1929, Dyschirius 229

aegeus J. Müller, 1921, Stenolophus 406

aegrota Bates, 1883, Odacantha [Casnonia] 444



aegrotus Bates, 1873, Trechus 325

aegyptia C. Zimmermann, 1832, Amara 552

aegyptiaca Dejean, 1831, Diplocheila [Rembus] 439

aegyptiaca Jedlika, 1960, Eucolliuris [Colliuris] 443

aegyptiaca Dejean, 1825, Myriochila [Cicindela] 114

aegyptiacum Dejean, 1831, Bembidion 251



aegyptiacus Motschulsky, 1844, Carabus 185

aegyptiacus Peyron, 1856, Cybister 50

aegyptiacus Schatzmayr, 1936, Dicheirotrichus 402



aegyptiacus Schatzmayr, 1936, Dyschirius 225

aegyptiacus Régimbart, 1883, Gyrinus 27

aegyptiacus Schatzmayr, 1936, Laemostenus 536

aegyptiacus Dejean, 1826, Licinus 441

aegyptiacus Dejean, 1828, Masoreus 357



aegyptiacus Mannerheim, 1837, Pheropsophus [Brachinus] 217

aegyptiacus Schatzmayr & Koch, 1934, Tachys [DA] 276



aegyptianus Mjöberg, 1905, Distichus [Scarites] 231

aegypticus Jedlika, 1952, Tetragonoderus 357



aegyptius Klug, 1832, Diplocheila [Rembus] 439

aegyptius Tschitschérine, 1897, Poecilus [Feronia] 479

aelleni Antoine, 1952, Laemostenus [Pristonychus] 539

aemiliae Vanni & Magrini, 1986, Duvalius 301

aemiliana A. Fiori, 1903, Amara 548

aemulus Fischer von Waldheim, 1823, Carabus 197

aemulus Putzeys, 1846, Dyschirius 228

aemulus Wollaston, 1864, Nesacinopus [Cratognathus] 388

aenea DeGeer, 1774, Amara [Carabus] 547



aenea Latreille, 1806, Loricera 98

aenea Gebler, 1825, Nebria 89

aeneicollis Grimmer, 1841, Poecilus 482

aeneicostis Bates, 1883, Brachinus 215



aeneipennis Sturm, 1815, Carabus 173

aeneipennis Faldermann, 1836, Harpalus [Omaseus] 371

aeneipes Bates, 1883, Bembidion 257

aeneo Boheman, 1849, Dyschirius 230



aeneobrachialis Bates, 1892, Dyschirius 230

aeneocephalus Dejean, 1826, Chlaenius 355



aeneocuprea Chaudoir, 1865, Cicindela 106

aeneocuprea Fairmaire, 1886, Pristosia [Calathus] 530

aeneocupreus G. Hauser, 1932, Carabus [Apotomopterus] 123

aeneocupreus Heinz, 1971, Lindrothius [Synuchidius] 529

aeneocupreus Fairmaire, 1887, Pterostichus [Platysma] 487

aeneola Poppius, 1906, Amara 547

aeneola Bates, 1873, Pristosia [Pristonychus] 530

aeneolus Chaudoir, 1876, Amblystomus [Hispalis] 360

aeneolus A. Morawitz, 1886, Carabus 178

aeneolus Reitter, 1900, Harpalus 371

aeneolus Chaudoir, 1868, Poecilus [Feronia] 479



aeneomicans Chaudoir, 1837, Amara [Bradytus] 552

aeneomicans Chaudoir, 1850, Apristus 431



aeneoopaca Motschulsky, 1860, Cicindela 106

aeneopiceus Stephens, 1828, Harpalus 23, 371

aeneopunctatus DeGeer, 1774, Carabus 160

aeneotinctum Bates, 1873, Agonum [Anchomenus] 454

aeneovirescens Straneo, 1939, Abacetus 346

aeneoviridimicans Netolitzky, 1938, Bembidion 267

aenescens Putzeys, 1866, Amara [Leiocnemis] 555



aenescens Depoli, 1929, Carabus 151

aenescens Casey, 1884, Harpalus 371

aenescens J. R. Sahlberg, 1880, Hydroporus 60

aenescens C. G. Thomson, 1870, Ilybius 41



aenescens Poppius, 1905, Ilybius [Agabus] 42

aenescens Chaudoir, 1850, Pterostichus [Feronia] 493

aeneum Germar, 1824, Bembidion 263

aeneum Motschulsky, 1859, Calosoma [Callisoma] 120

aeneuoides J. Jacquet, 1942, Harpalus 376

aeneus Dejean, 1828, Abacetus [Feronia] 346

aeneus Putzeys, 1873, Calathus 527

aeneus Motschulsky, 1850, Carabus 186

aeneus Motschulsky, 1865, Catascopus 435

aeneus Fischer von Waldheim, 1823, Cychrus 19, 202

aeneus Dejean, 1825, Dyschirius [Clivina] 224

aeneus Stephens, 1828, Gyrinus 27


aeneus Fabricius, 1775, Harpalus [Carabus] 371

aeneus Panzer, 1796, Ilybius [Dytiscus] 42

aeneus Dejean, 1828, Poecilus [Feronia] 483

aeneus Fischer von Waldheim, 1829, Pterostichus [ND] 514



aeneus Solsky, 1871, Pterostichus [Steropus] 493

aenigma Chaudoir, 1869, Abacetus 346

aenigma Tschitschérine, 1897, Harpalus [Ophonus] 384

aenigmaticus Uéno, 1978, Yamautidius 345

aequabilis Gschwendtner, 1923, Agabus [Gaurodytes] 35

aequale A. Morawitz, 1863, Omophron 208

aequaliceps Reitter, 1896, Carabus [Cechenus] 137

aequalis Sharp, 1882, Agabus 35

aequalis Jeannel, 1931, Duvalius 306

aequalis A. Zimmermann, 1918, Graptodytes [Hydroporus] 59

aequalis Falkenström, 1932, Hygrotus 73

aequatorialis Baliani, 1939, Amara 550

aequatorius A. Zimmermann, 1917, Cybister 50

aequatorius Guignot, 1936, Synchortus 34

aequatum Jedlika, 1936, Agonum [Colpodes] 454



aequatus Aubé, 1836, Brychius 30

aequatus Motschulsky, 1844, Carabus 183

aequatus Audinet‑Serville, 1821, Licinus 441



aequicollis Motschulsky, 1866, Abacetus [Astygis] 346

aequicollis Motschulsky, 1844, Harpalus 371

aequicostalis Staven & Heinz, 1995, Carabus 193

aequicostatus Mandl, 1974, Carabus 162

aequilimes Semenov, 1898, Carabus 190

aequimarginatus Falkenström, 1936, Rhantus 47

aequistriatus Kraatz, 1878, Carabus 173

aequistriatus Kurnakov, 1961, Lindrothius 529

aerarius Coquerel, 1859, Poecilus [Feronia] 481

aerata Stephens, 1828, Amara 549

aeratus Stephens, 1828, Agabus [Colymbetes] 37

aeratus Géhin, 1885, Carabus 135

aeratus Quensel, 1806, Chlaenius [Carabus] 355



aeratus Stephens, 1828, Dyschirius 224

aeratus Stephens, 1835, Gyrinus 27



aeratus Hope, 1831, Nirmala [Omaseus] 476

aeratus Chaudoir, 1861, Peronomerus 448

aerea Chevrolat, 1841, Calomera [Cicindela] 100

aereicollis G. Hauser, 1921, Carabus [Acoptolabrus] 122

aereipennis Solsky, 1871, Pterostichus [Steropus.] 493

aereum Jacquelin du Val, 1851, Bembidion 271

aereus Motschulsky, 1864, Apristus 431

aereus Dejean, 1826, Carabus 135


aereus Fischer von Waldheim, 1820, Carabus 169

aereus Klug, 1834, Dineutus [Gyrinus] 26



aereus Ahrens, 1830, Dyschirius [Clivina] 224

aeripennis Andrewes, 1931, Holcoderus 436



aerosum Erichson, 1837, Bembidion 266

aerosus Coiffait, 1962, Aphoenops 294



aeruginosa Kolenati, 1845, Amara 548

aeruginosiformis Breuning, 1932, Carabus 174

aeruginosum Gebler, 1833, Bembidion [Peryphus] 265

aeruginosus Fischer von Waldheim, 1820, Carabus 169



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