Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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brevepilosa Antoine, 1941, Cymindis 411

brevicaudis Belousov & Kabak, 1993, Trechus 328

brevicaudus Imura & Mizusawa, 1995, Carabus 148

breviceps Bates, 1883, Dromius 420

brevicolIis Joseph, 1870, Laemostenus [Sphodrus] 535

brevicollis Baudi di Selve, 1882, Acinopus 368

brevicollis Chaudoir, 1850, Amara [Leirus] 559

brevicollis Chaudoir, 1844, Anisodactylus 360

brevicollis Reiche, 1875, Badister 440

brevicollis Motschulsky, 1844, Brachinus 213

brevicollis Lapouge, 1910, Carabus 141

brevicollis Chaudoir, 1843, Chlaenius [Epomis] 353

brevicollis Dejean, 1826, Licinus 442

brevicollis Jeannel, 1960, Nannotrechus 315

brevicollis Fabricius, 1792, Nebria [Carabus] 92

brevicollis Audinet‑Serville, 1821, Ophonus [Harpalus] 389



brevicollis Reitter, 1884, Penthus 367

brevicollis Reiche, 1862, Platyderus 522

brevicollis Mannerheim, 1825, Pogonus 287

brevicollis Putzeys, 1866, Sparostes 221

brevicollis N. Ito, 1998, Trichotichnus 394

brevicollis Schaum, 1857, Zabrus 570



brevicornis Ménétriés, 1851, Amara [Leirus] 558

brevicornis Kraatz, 1878, Carabus 180

brevicornis Germar, 1824, Harpalus 373

brevicornis Uéno, 1979, Kurasawatrechus 313

brevicornis Heller, 1903, Lioptera 436

brevicornis Chaudoir, 1846, Polyderis [Tachys] 274

brevicornis Gautier des Cottes, 1869, Pterostichus [Omaseus] [HN] 509

brevicornis Kirby, 1837, Pterostichus [Argutor] 490

brevicornis Jeannel, 1930, Thalassophilus 346

brevicornutus Luna de Carvalho, 1973, Paussus 210

brevicorpus Belousov & Kabak, 1993, Trechus 328

breviculus Cavazzuti & Ratti, 1998, Carabus 190

breviformis Chaudoir, 1863, Carabus 191

breviformis Tschitschérine, 1898, Microderes [Neopangus] 387

breviformis Mlyná, 1977, Molops 474


brevilabris Kirby, 1837, Amara [Curtonotus] 561

brevilama Straneo, 1989, Pterostichus 510

brevilimbata Pic, 1922, Lebia [DA] 428

brevinotum Morvan, 1973, Bembidion [Nepha] 253

brevior Poppius, 1907, Carabus 133

brevior Nakane, 1963, Haliplus 31

brevior Uéno, 1980, Trechiama 322



brevipenis Mateu, 1950, Pterostichus [Haptoderus] 492

brevipennis Casey, 1924, Amara [Curtonotus] 561

brevipennis Chaudoir, 1844, Amara [Bradytus] 553

brevipennis Lapouge, 1913, Carabus 138

brevipennis J. Frivaldszky, 1879, Dicropterus [Scotodipnus] 239



brevipennis Schauberger, 1921, Molops 474

brevipennis Winkler, 1926, Neotrechus 315

brevipennis Chaudoir, 1850, Pachycarus [Pachycarus] 367

brevipennis Ménétriés, 1848, Polistichus 574

brevipennis Semenov, 1889, Pseudotaphoxenus 540

brevipennis Chevrolat, 1840, Pterostichus [Omaseus] 496

brevipennis Wollaston, 1864, Syntomus [Metabletus] 432

brevipilosa W. Horn, 1908, Calomera [Cicindela] 99

brevipilosus Knirsch, 1927, Duvalius [Duvalites] 298



brevis Aubé, 1837, Acilius 49

brevis Dejean, 1828, Amara 554

brevis Gautier des Cottes, 1866, Calathus 524

brevis Dejean, 1826, Carabus 167

brevis Aubé, 1838, Cybister 50

brevis W. Horn, 1905, Cylindera [Cicindela] 110

brevis Uéno, 1978, Epaphiopsis 308

brevis Wehncke, 1880, Haliplus 31

brevis Motschulsky, 1844, Harpalus 373

brevis R. F. Sahlberg, 1834, Hydroporus 60

brevis Sturm, 1835, Nebrioporus [Hydroporus] 66

brevis G. R. Waterhouse, 1833, Notiophilus 97

brevis Duftschmid, 1812, Pterostichus [Carabus] 506



brevis Faldermann, 1836, Pterostichus [Steropus] 493

brevis Lindroth, 1956, Synuchus 545



brevis Jedlika, 1958, Trichotichnus 395

brevistriga Régimbart, 1906, Bidessus 55

brevistylis Jeannel, 1942, Pterostichus [Platysma] 499

brevitarsis Normand, 1946, Bradycellus 400

brevitarsis Normand, 1933, Cymindis 414

brevithorax Machado, 1992, Calathidius 532

brevituberculatus Roubal, 1909, Carabus 196

brevius Netolitzky, 1918, Bembidion 250



breviuscula V. E. Jakovlev, 1887, Cymindis 417

breviusculum Mannerheim, 1830, Calosoma [Carabus] 118

breviusculus Kraatz, 1876, Carabus 185

breviusculus Kraatz, 1887, Carabus 179

breviusculus Chaudoir, 1846, Harpalus 374

breviusculus Poppius, 1905, Hydroporus 60

breviusculus Gschwendtner, 1935, Laccornis 75

breviusculus Solsky, 1873, Notiophilus 96

breviusculus R. F. Sahlberg, 1844, Pterostichus [Feronia] 490

breviusculus Mannerheim, 1849, Synuchus [Taphria] 545



breviusculus K. Daniel & J. Daniel, 1906, Trechus 328

brevivittis Bedel, 1907, Glycia 409

brezinai Deuve, 1994, Carabus 146

brezinai Deuve, 1993, Cychrus 203

brezinai Dvoák, 1994, Leistus 81

brezinai Deuve & Quéinnec, 1992, Trechus 23, 328



brezinaorum Deuve & Quéinnec, 1992, Trechus 338

briali Antoine, 1934, Amara 562

brianii Mancini, 1912, Duvalius [Anophthalmus] 302



brianskyi Lutshnik, 1909, Carabus 132

brincki Guignot, 1946, Hyphydrus 74

brisouti Csiki, 1929, Amara 568

brisouti Abeille de Perrin, 1876, Hydraphaenops [Trechus] 311

brisouti Holdhaus, 1912, Microlestes 422

brisouti Bedel, 1895, Reicheia 222

brisoutii Fauvel, 1863, Carabus 168

britannicus Born, 1908, Carabus 166

brittoni Fassati, 1955, Bembidion 260

brittoni Ali, 1967, Brachinus 215

brittoni Mandl, 1965, Carabus 162

brittoni Mateu, 1957, Dromius 420

brittoni Basilewsky, 1954, Hystrichopus 418

brittoni Quentin, 1952, Mastax 216

brittoni Basilewsky, 1948, Metadromius [Philorhizus] 421

brittoni Reichensperger, 1957, Paussus 210

brittoni Mateu, 1956, Pseudomesolestes [Mesolestes] 426

brittoni Habu, 1958, Pterostichus 513



brittoni Habu, 1955, Synuchus [Calathus] 546

brivioi Sciaky, 1998, Stomis 517

brockmanni Kleinfeld & Schütze, 1997, Carabus 142

brockmannioides Deuve & Mourzine, 1997, Carabus 142

broderi Daffner, 1992, Anophthalmus 291

brodskii Jankowski, 1937, Cicindela 106

brondelii Reiche, 1872, Zabrus 570

brosciformis Semenov, 1897, Carabus 147



brosseti Cabidoche, 1962, Laemostenus [Ceuthosphodrus] 535

brostenensis E. Jacquet, 1886, Carabus 185

browneanus Sharp, 1913, Haliplus 31

browneanus Guignot, 1959, Hydrovatus 71

brownei Sharp, 1913, Haliplus 31

brownei Guignot, 1949, Nebrioporus [Potamonectes] 66

browni Kamiya, 1934, Agabus 35

browni Leech, 1938, Agabus 37

browni Deuve, 1999, Carabus 166

broziki Hrka & Pulpán, 1980, Duvalius 306

brucei Andrewes, 1923, Amara 558

brucei Andrewes, 1930, Pseudotaphoxenus [Taphoxenus] 540

brucki Piccioli, 1870, Duvalius [Anophthalmus] 302

brucki Wehncke, 1875, Hydroporus 60

bruckii Schaum, 1859, Pterostichus 511

bruckii Fairmaire, 1862, Trechus 328



bructeri Panzer, 1796, Leistus [Carabus] 83

brueggemanni Verhoeff, 1891, Agonum [Anchomenus] 451

bruggei Gottwald, 1990, Carabus 123

bruggei Deuve, 1991, Cychrus 203

bruggeianus Deuve, 1992, Carabus 175

brujasi Sainte‑Claire Deville, 1901, Duvalius [Trechus] 302



brullei Géhin, 1885, Carabus 194

brullei Chaudoir, 1874, Myas 475

brumalis Casey, 1918, Amara [Celia] 564

bruneti Jeannel, 1926, Aphoenops 294

brunicolle J. Schmidt, 1995, Xestagonum 467

brunii Gredler, 1873, Pterostichus [Feronia] 488

brunnea Gyllenhal, 1810, Amara [Harpalus] 556

brunnea Putchkov, 1993, Cephalota [Cicindela] 102

brunnea Jedlika, 1951, Formosiella 436



brunnea Chaudoir, 1861, Hexagonia 407

brunnea Duftschmid, 1812, Oreonebria [Carabus] 95



brunneicollis J. R. Sahlberg, 1913, Tachyura [Tachys] 279

brunneicupreus Deuve, 1994, Carabus 193



brunneipennis A. Costa, 1847, Percus 478

brunneipennis Straneo, 1955, Pterostichus 501



brunneiventris Louwerens, 1949, Styphlomerus 218

brunneomarginatus N. Ito, 1998, Trichotichnus 394



brunnescens Casey, 1918, Amara [Celia] 564

brunnescens Kirschenhofer, 1988, Apristus 431



brunnescens Motschulsky, 1861, Clivina [Eupalamus] 220

brunneus Fabricius, 1798, Agabus [Dytiscus] 39



brunneus Kamiya, 1935, Agabus 35

brunneus Brullé, 1832, Calathus 524

brunneus Dejean, 1831, Dicrodontus [Polystichus] 573



brunneus Hope, 1831, Laemostenus [Sphodrus] 537

brunneus Karsch, 1881, Platyderus 521

brunneus Sturm, 1824, Pterostichus [Platysma] [NO] 511

brunneus Uéno, 1977, Stevensius 320

brunneus N. Ito, 1998, Trichotichnus 394

brunnicorne Dejean, 1831, Bembidion 258



brunnicornis Heer, 1837, Amara 548

brunnicornis Motschulsky, 1864, Brachinus 214

brunnipennis Dejean, 1831, Amara 558

brunnipes Sturm, 1825, Acupalpus [Trechus] 397



brunnipes C. R. Sahlberg, 1827, Bembidion 271

brunnipes Dejean, 1831, Bembidion 268

brunnipes Sturm, 1825, Bembidion 259

brunnipes Beuthin, 1885, Carabus [Platychrus] 185

brunnipes Fischer von Waldheim, 1829, Harpalus 386



brunnipes Dejean, 1828, Molops 473

brunnipes Stephens, 1828, Pterostichus 512

brunnipes Dejean, 1825, Siagona 218

brunniventris Motschulsky, 1864, Brachinus [ND] 215

bruschii Tarrier, 1974, Carabus 166

bruschii Vigna Taglianti, 1999, Duvalius 302

bruskelchus Morvan, 1996, Platynus [Batenus] 462

bruskornekum Morvan, 1996, Xestagonum 467

brussanus Straneo, 1986, Tapinopterus 518

brutius Straneo, 1935, Poecilus [Pterostichus] 482



bruttianus Born, 1906, Carabus 161

bruxellense Wesmael, 1835, Bembidion 260



bryanti Carr, 1930, Agabus 36

bryanti Carr, 1932, Bembidion 247

brzotinensis Janák, 1987, Duvalius 307

bualei Jacquelin du Val, 1852, Bembidion 260

bucciarelli Sciaky, 1979, Anisodactylus 361

bucciarellii Monguzzi, 1976, Badister [Baudia] 440

bucciarellii Monguzzi, 1982, Boldoriella 296



bucciarellii Mandl, 1966, Carabus 197

bucciarellii Tamanini, 1954, Orotrechus 317

bucciarellii Pesarini, 1979, Stomis 517

bucculentus Kraatz, 1874, Harpalus 382

bucculentus Andrewes, 1935, Platytarus 410



bucculentus Reitter, 1899, Scarites 233

bucephalum Netolitzky, 1920, Bembidion 258



bucephalus Dejean, 1829, Acinopus 369

bucephalus Dieck, 1869, Aphoenops [Anophthalmus] 294



bucephalus Dieck, 1870, Aphoenops [Anophthalmus] 294

bucephalus Olivier, 1795, Dixus [Scarites] 365

bucephalus Dejean, 1828, Molops [Feronia] 472

bucerdo Ménétriés, 1832, Scarites 234

buchannani Hope, 1831, Metacolpodes [Colpodes] 460

bucharica Tschitschérine, 1898, Amara 566

bucharicum Netolitzky, 1934, Bembidion 268

bucharicus Liebke, 1928, Brachinus 216

bucharicus Tschitschérine, 1898, Harpalus 374

bucharicus Jedlika, 1961, Pseudotaphoxenus [Taphoxenus] 541

buchariplaga Netolitzky, 1943, Bembidion 249



bucheti J. Ochs, 1949, Carabus [Eutelocarabus] 191

bucheti Alluaud, 1919, Licinopsis 540

bucheti Régimbart, 1898, Nebrioporus [Hydroporus] 66



bucheti J. Ochs, 1937, Sphodropsis 543

buchi G. Hauser, 1913, Carabus [Coptolabrus] 143

buchtarmensis Kryzhanovskij, 1953, Carabus 199

bucida Pallas, 1776, Scarites [Carabus] 233



bucidus Reiche & Saulcy, 1855, Odotoncarus [Ditomus] 366

bucurensis Janák, 1993, Duvalius 306

budae Kenderessy, 1879, Duvalius [Anophthalmus] 298

buddaicus buddaicus Semenov, 1887, Carabus 190



budensis Csiki, 1946, Carabus 197

budensis Csiki, 1908, Cymindis 412

budha Breuning, 1950, Carabus 146

budhaica Deuve, 1988, Epaphiopsis [Epaphiama] 308

buettikeri Mateu, 1990, Dromius 420

buettikeri Mateu, 1986, Merizomena [Glycia] 409

buettikeri Brancucci, 1985, Yola 57

buffi Deuve & Kaláb, 1992, Carabus 145

bufo Sharp, 1890, Clypeodytes [Bidessus] 55

bufo Fabricius, 1801, Platytarus [Galerita] 410

buglanensis Bulirsch, 1996, Dyschirius 228

buglaniensis Kirschenhofer, 1981, Pterostichus 500

bugnioni K. Daniel, 1902, Bembidion 245

bugnionii Chaudoir, 1837, Carabus 184

bukoveci Pretner, 1949, Anophthalmus 291

bulgani Jedlika, 1969, Agonum 454

bulgani Jedlika, 1968, Bembidion 248

bulgani Jedlika, 1968, Pogonus 287

bulgardagense Fassati, 1990, Bembidion 259



bulgaricus Csiki, 1943, Agabus [Gaurodytes] 39

bulgaricus Csiki, 1927, Carabus 129

bulgaricus Knirsch, 1924, Duvalius [Trechus] 308



bulgaricus Hlisnikovský, 1955, Hydroporus 60

bulgaricus Maan, 1938, Molops 473



bulgaricus Roubal, 1928, Patrobus 285

bulgaricus Lutshnik, 1915, Pterostichus [Platysma] 499

bulgaricus Obenberger, 1917, Pterostichus 509

bulgaricus Pawowski, 1972, Trechus 22, 333

bulgarus Csiki, 1930, Pterostichus 509

bulghardaghensis Breuning, 1926, Carabus 130



bulgharensis Jedlika, 1958, Harpalus 382

bulgharmaadensis E. Bodemeyer, 1915, Carabus [Procerus] 185



bulimus Motschulsky, 1850, Scarites 233

bulirschi Dvoák, 1995, Laemostenus 534

bullaquensis Zaballos & Ruiz‑Tapiador, 1997, Typhlocharis 241

bullata Marseul, 1871, Amara 552

bullatus Schrank, 1798, Badister [Carabus] 440



bulsanense Gredler, 1863, Bembidion 249

bulwerii Stephens, 1828, Pterostichus [Omaseus] 487

bungei Zaitzev, 1910, Hydroporus 63

bungei Tschitschérine, 1894, Pterostichus [Feronia] 505

bungii Gebler, 1829, Carabus 191

bungii Chaudoir, 1844, Harpalus 374

buparius Forster, 1771, Scarites [Tenebrio] 233

buqueti Guérin‑Méneville, 1838, Siagona 218

burdigalensis Schuler, 1964, Amara [Celia] 557

bureianus Shilenkov, 1996, Carabus 165

bureli Morvan, 1994, Pterostichus 510

bureli Morvan, 1995, Xestagonum [Colpodes] 467

bureschi Jeannel, 1928, Duvalius 308

bureschi Maan, 1933, Tapinopterus 519

bureschianus Breuning, 1928, Carabus 185



buresi Hanuš, 1923, Carabus [Promorphocarabus] 174

buresi Maan, 1938, Hydroporus 63

buresi Knirsch, 1923, Pheggomisetes [Trechus] 318



burgassiensis Apfelbeck, 1904, Carabus 131

burghauseri A. Fleischer, 1899, Carabus 173

buriatus Lapouge, 1925, Carabus 171

burjaticus Poppius, 1906, Pterostichus [Cryobius] 491

burmeisteri Fischer von Waldheim, 1842, Cicindela 105

burmeisteri Gistel, 1837, Cicindela 105

burmeisteri Heer, 1838, Pterostichus 488



burnaschewi Dejean, 1826, Carabus 170

burrellii Curtis, 1824, Pogonus 287

buschi Semenov & Znojko, 1932, Carabus 137

bushirica Jedlika, 1946, Cymindis 412

busii F. Battoni, 1984, Calathus 524

busii Magrini & Vanni, 1984, Duvalius 304

businskae Dvoák, 1998, Broscosoma 235

businskyi Lassalle, 1999, Acalathus [Procalathus] 529


businskyi Deuve, 1990, Carabus 154

businskyi Deuve, 1992, Cychropsis 201

businskyi Dvoák, 1994, Leistus 81

businskyorum Imura, 1996, Carabus 133

businskyorum Ledoux & Roux, 1997, Nebria 90

bussleri Fery, 1994, Graptodytes 59

buttonii Straneo, 1959, Percus 478

buycki G. Hauser, 1924, Carabus [Apotomopterus] 124

bythiniensis Maan, 1944, Pterostichus [Haptotapinus] 496

bytinskii Schweiger, 1970, Carabus 188

byzantinus Casale, M. Etonti & Giachino, 1992, Anillidius [Troglocimmerites] 290



byzantinus Apfelbeck, 1904, Licinus 442

byzantinus Apfelbeck, 1902, Molops 474

byzantinus Ganglbauer, 1897 (as 1896), Tapinopterus 519

byzantinus Apfelbeck, 1901, Trechus 328

bzybicus Belousov, 1998, Nannotrechus 315

bzybiensis Gottwald, 1982, Carabus 201

cabidochei Coiffait, 1959, Aphoenops 293

cabidochei Colas, 1966, Pterostichus [Haptoderus] 489

cabrerai Enderlein, 1929, Carabus 176

cabrerai Jeannel, 1936, Trechus [Anchotrechus] 328

cabrerensis Mateu, 1972, Pterostichus 503

cabuljensis Apfelbeck, 1919, Carabus 163

cabuljensis Mlyná, 1977, Molops 473

cabuljensis Winkler, 1933, Omphreus 445

cadamuroi Tamanini, 1954, Orotrechus 317

cadevalli Ferrer y Dalmau, 1911, Cicindela 108

cadilhaci Antoine, 1951, Laemostenus [Pristonychus] 537

cadouali Morvan, 1977, Amara 566

cadoudali Morvan, 1976, Trechus 329

caducum Casey, 1918, Bembidion 262

cadurcus Jeannel, 1955, Duvalius 302

caeculus Belousov & Kabak, 1993, Aepiblemus 289

caecus Jacquelin du Val, 1851, Anillus 238

caecus Antoine, 1941, Mesolestes [Metabletus] 421

caecus Uéno & Yoshida, 1966, Oroblemus 316

caecus Jeannel, 1938, Thalassophilus 346

caelatus Fabricius, 1801, Carabus 163



caelatus Weber, 1801, Chlaenius [Tachypus] 348

caerulans A. Morawitz, 1886, Carabus 153

caerulans Andrewes, 1938, Euschizomerus 447

caeruleatus Bates, 1878, Harpalus 374

caeruleiceps Bates, 1892, Callistoides [Chlaenius] 347

caeruleum Krynicki, 1832, Bembidion 247

caeruleus Panzer, 1810, Carabus 173


caeruleus Voet, 1778, Carabus [Buprestis] 166

caesareae Casale, 1988, Laemostenus 538

caesareum Netolitzky, 1914, Bembidion 255

caesareus Semenov, 1906, Carabus 165

caesus Duftschmid, 1805, Peltodytes [Dytiscus] 33

caffer Duftschmid, 1812, Harpalus [Carabus] 386

cailloli Sainte‑Claire Deville, 1902, Duvalius [Trechus] 302

caiphus Reiche & Saulcy, 1855, Harpalus 374

caja Samko, 1938, Cicindela 107

cakorensis Mlyná, 1977, Molops 472



calabrica Mandl, 1944, Cicindela 103

calabricus Guignot, 1957, Bidessus 55



calabrus Leoni, 1908, Calathus 525

calabrus A. Fiori, 1903, Carabus 179

calabrus Schauberger, 1926, Ophonus [Harpalus] 392

calabrus Flach, 1907, Poecilus 482

calandrii Casale & Vigna Taglianti, 1990, Duvalius 302

calashensis Deuve, 1982, Trechus 328

calathiformis Wollaston, 1865, Gomerina [Pterostichus] 521

calathiformis N. Ito, 1999, Trichotichnus 394

calathinus Lindroth, 1956, Synuchus 545

calathoides Putzeys, 1866, Amara [Leirides] 562


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