Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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chinensis Bousquet & Smetana, 1996, Paropisthius 98

chinensis Jedlika, 1960, Peliocypas [Risophilus] 434

chinensis Jedlika, 1963, Pheropsophus 217

chinensis Jedlika, 1934, Platynus [Colpodes] 464

chinensis Jedlika, 1962, Poecilus [Pterostichus] 481

chinensis Habu, 1986, Pogonistes 286

chinensis Jedlika, 1933, Pristosia [Kanoldia] 531

chinensis Jedlika, 1937, Pristosia [Eucalathus] 24, 531

chinensis Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 540



chinensis Falkenström, 1936, Rhantus 47

chinensis Erichson, 1834, Scarites 234

chinensis Jedlika, 1962, Steropanus [Tritrichis] 517

chinensis Jedlika, 1932, Stomis 517

chinensis Lindroth, 1956, Synuchus 545



chinensis Jedlika, 1932, Tachyura [Tachys] 279

chinensis Jeannel, 1920, Trechus 325



chinensis Csiki, 1907, Trichisia 448

chinensis Fairmaire, 1886, Trichotichnus [Zabrus] 394



chingana Baliani, 1939, Amara 559

chinganensis Semenov, 1898, Carabus 144

chinkiangensis Kirschenhofer, 1991, Pterostichus 505



chionobatus Born, 1918, Carabus 138

chionobatus Apfelbeck, 1908, Trechus 342

chionobatus Apfelbeck, 1912, Trechus 342

chionophilus Apfelbeck, 1904, Carabus 198

chiosanus Reitter, 1889, Zabrus 570

chiponica Jedlika, 1939, Lebia 428

chiragricus Fischer von Waldheim, 1828, Carabus 192

chiricola Kwon & Lee, 1984, Carabus [Damaster] 122

chiricus Lassalle, 1999, Carabus 159

chirisanensis Kwon & Lee, 1984, Carabus 183

chishimanus Nakane, 1961, Carabus [Procrustes] 164

chishimanus Kôno, 1944, Ilybius 42

chitral Andrewes, 1935, Bembidion 260

chitralensis Shilenkov & Heinz, 1988, Nebria 90

chitralicus Casale, 1988, Laemostenus [Pristonychus] 537

chitwanae Acciavatti & Pearson, 1989, Cosmodela [Cicindela] 108



chivense Lutshnik, 1934, Agonum 453

chivensis Lutshnik, 1928, Amara 551

chivensis Fedorenko, 1992, Dyschirius 226



chlaenioides Gistel, 1857, Anisodactylus 361

chlaenioides LaFerté‑Sénectère, 1851, Chlaenius [Amblygenius] 349

chloe Dokhtouroff, 1887, Jansenia [Thopeutica] 112

chloodes Andrewes, 1935, Chlaenius 352

chloreis Bates, 1883, Agonum [Colpodes] 454

chloreum Bates, 1873, Bembidion 243

chloris Hope, 1831, Calomera [Cicindela] 100



chloris Dejean, 1831, Cicindela 105

chloriventris Motschulsky, 1862, Lebia [Lebida] 427

chlorizans Bates, 1892, Coryza [Clivina] 221

chlorizans Bates, 1883, Harpalus 374

chlorizans Solsky, 1874, Ophonus 388

chlorocephala J. J. Hoffmann, 1803, Lebia [Carabus] 427

chlorocephalus Kollar, 1836, Pristomachaerus [Panagaeus] 356



chlorochila Chaudoir, 1852, Cylindera [Cicindela] 112

chlorodius Dejean, 1826, Chlaenius 349



chlorophanum Sturm, 1825, Bembidion 266

chlorophanus Panzer, 1799, Ophonus [Carabus] 388

chloropleura Chaudoir, 1865, Jansenia [Cicindela] 112



chloropterus Stephens, 1828, Harpalus 381

chloropus Bates, 1883, Bembidion 257



chloropus Audouin & Brullé, 1839, Cylindera [Cicindela] 110

chlorostictum Dejean, 1831, Calosoma 120

chlorotica Fairmaire, 1867, Amara 567

chloroticus Fischer von Waldheim, 1823, Cardiaderus [Daptus] 286

chloroticus Dejean, 1829, Dicheirotrichus [Harpalus] 402

chloroticus Régimbart, 1887, Laccophilus 76

chlorotrachelus G. Hauser, 1921, Carabus [Coptolabrus] 141

chobauli Jeannel, 1942, Microlestes 423



chobauti Puel, 1926, Acinopus 23, 368

chobauti Puel, 1924, Amara 549

chobauti Puel, 1924, Dromius 420



chobauti Reitter, 1900, Harpalus 375

chobauti Sainte‑Claire Deville, 1905, Harpalus [Licinoderus] 374

chobautianus Lutshnik, 1922, Harpalus 374

chodjaii Morvan, 1975, Amara 566

chodjaii Morvan, 1975, Cymindis 412

chodjaii Morvan, 1974, Platyderus 522

chodjaii Morvan, 1974, Trechus 328

chodshenticus Ballion, 1871, Eocarterus [Ditomus] 365

chogyesanus Park, Kwon & Lafer, 1996, Pterostichus 499

chokyuryongensis Imura & Yamaya, 1994, Carabus 159

cholaensis Deuve, 1996, Trechus 328

cholashanus Imura, 1998, Carabus 176

chomolungma Deuve & Quéinnec, 1985, Trechus 328



chonmanus Imura & Yamaya, 1994, Carabus 199

chonmasanensis Deuve, 1994, Carabus 122

chormaensis Hieke, 1995, Amara 559

chormaensis Deuve, 1994, Carabus 178

chormaensis Deuve, 1993, Trechus 328


chotanensis Tschitschérine, 1893, Pterostichus [Feronia] 509

chotaroi Imura, Dejima & Mizusawa, 1993, Carabus 177

chotjaii Morvan, 1973, Agonum 454

chotjaii Morvan, 1973, Bembidion [Bembidionetolitzkya] 245

choui Deuve, 1989, Carabus 190

christinae Merceron, 1998, Pterostichus 512

christophi Kraatz, 1878, Carabus 174

chrysis Reitter, 1892, Lebia 427

chrysites Bates, 1892, Trigonotoma 520

chrysitis Motschulsky, 1839, Carabus 192

chrysocephala Motschulsky, 1864, Lebia [Lamprias] 427

chrysocephalus P. Rossi, 1790, Chlaenius [Carabus] 355



chrysochlorus Fischer von Waldheim, 1812, Carabus 198

chrysomyia Bates, 1888, Lebia 428

chrysopus Reitter, 1887, Harpalus 374

chrysostomus Gozis, 1914, Hydroporus 65

chrysothorax Kraatz, 1886, Carabus 171

chrysothorax Krynitzky, 1832, Chlaenius 350

chugokuensis Nakane, 1961, Carabus [Apotomopterus] 177

chugokuensis N. Ito, 1996, Trichotichnus 395

chui Uéno, 1990, Trechiama 322

chuji Jedlika, 1956, Agonum 454

chuji Jedlika, 1956, Brachinus 212

chuji Jedlika, 1946, Chlaenius 353

chuji Jedlika, 1951, Sofota 430

chuji Jedlika, 1949, Trichotichnus 395

chujianum Jedlika, 1956, Xestagonum [Colpodes] 469

chujoi Jedlika, 1962, Agonum [Colpodes] 454

chujoi Jedlika, 1951, Bembidion 267



chujoi Jedlika, 1956, Brachinus 216

chujoi Habu, 1960, Calleida 408

chujoi Habu, 1975, Chydaeus 362

chujoi Habu, 1973, Gnathaphanus 362

chujoi Satô, 1982, Platynectes 44

chujoi Habu, 1959, Pterostichus 507

chujoi Habu, 1961, Trephionus 546



chujoianum Jedlika, 1962, Bembidion 254

chujoianus Jedlika, 1962, Pterostichus 498

chujoiellus Jedlika, 1962, Pterostichus 511

chulii J. Schmidt, 1995, Deltomerodes 281

chunghusorum Lutshnik, 1922, Pterostichus [Platysma] 486

chungkingi Jedlika, 1932, Pterostichus 500

chusanensis A. Nicolas, 1924, Carabus 156

chusyae Machado, 1987, Canarobius 345

chydaeus Tschitschérine, 1897, Pterostichus [Feronia] 496


cicatricosoides Breuning, 1961, Carabus 197

cicatricosoides Mandl, 1955, Carabus 145

cicatricosulus A. Morawitz, 1886, Carabus 192

cicatricosus Fischer von Waldheim, 1842, Carabus 145

cicatricosus Motschulsky, 1865, Chlaenius [Epomis] 353

cicioarae Jeannel, 1930, Duvalius 299



ciffrei Barbier, 1885, Stenolophus 406

ciftekopruleriensis P. Machard, 1975, Carabus 181

ciliatum Vauloger de Beaupré, 1898, Zuphium 574

ciliatus Liebke, 1933, Brachinus 215

ciliatus Wollaston, 1862, Calathus 526

ciliatus Olivier, 1791, Ilybius [Dytiscus] 42

ciliatus Uéno, 1955, Ryugadous 319

cilicicum De Monte, 1947, Bembidion 260

cilicicus Csiki, 1932, Ophonus [Harpalus] 389

ciliciensis Mlyná, 1979, Harpalus 381

ciliciensis Schauberger, 1927, Ophonus [Harpalus] 389

cilicium Peyron, 1858, Zuphium 574

cilicius Straneo, 1986, Tapinopterus 518

cilihumerus Huang, Hu & Sun, 1994, Harpalus 384

ciloensis Deuve, 1994, Carabus 147

cimensis Cockerell, 1922, Nesarpalus [Harpalus] 24, 388

cimensis Cockerell, 1922, Scarites 233

cimicoides O. F. Müller, 1776, Laccophilus [Dyticus] 76

cimmerium Andrewes, 1922, Bembidion 245

cimmerius Fischer von Waldheim, 1823, Laemostenus [Sphodrus] 537

cincta Brullé, 1839, Cymindis 412



cincta Ledoux & Roux, 1991, Nebria 91

cincta Straneo, 1983, Nirmala 476

cinctella Andrewes, 1925, Nebria 90



cinctellum Chevrolat, 1882, Callytron [Cicindela] 101

cinctellus Chaudoir, 1876, Brachinus 215



cincticollis Mäklin, 1881, Ilybius 42

cincticollis Chevrolat, 1840, Platyderus [Argutor] 523

cinctipennis Boheman, 1848, Chlaenius 351

cinctus Motschulsky, 1850, Calathus 527



cinctus O. F. Müller, 1776, Hydaticus [Dyticus] 54

cinctus Sharp, 1882, Ilybius 42

cinctus Motschulsky, 1860, Patrobus 21, 285

cinereus Linné, 1758, Graphoderus [Dytiscus] 49



cinereus Aubé, 1836, Haliplus 32

cingulata Dejean, 1825, Cymindis 412

cingulatus Gebler, 1843, Agatus [Dromius] 24, 408

cingulifer Guignot, 1956, Laccophilus 76

circassica Reitter, 1888, Amara 560


circassicum Reitter, 1890, Bembidion [Pseudolimnaeum] 256

circassicus Ganglbauer, 1886, Carabus [Tribax] 200

circassicus Reitter, 1890, Deltomerus 282

circassicus Reitter, 1888, Porocimmerites [Anophthalmus] 318



circassius Reitter, 1887, Laemostenus [Antisphodrus] 533

circe Cavazzuti & Ratti, 1998, Carabus 124



circovichi Schatzmayr, 1944, Molops 472

circovichi J. Müller, 1922, Typhlotrechus [Trechus] 344



circularis Baliani, 1937, Amara 558

circularis Régimbart, 1895, Hyphydrus 74

circumcinctus Reiche, 1862, Acupalpus 398

circumcinctus R. F. Sahlberg, 1834, Bradycellus [Acupalpus] 401

circumcinctus Ahrens, 1811, Dytiscus 51

circumcinctus Motschulsky, 1858, Loxoncus [Megrammus] 404

circumdata Baliani, 1937, Amara 567

circumdata Dejean, 1822, Cephalota [Cicindela] 102



circumdata Krynicki, 1832, Cephalota [Cicindela] [HN] 102

circumdata Shibata, 1987, Parena 433

circumdatus Brullé, 1835, Chlaenius 351

circumducta Reitter, 1890, Lebia [Glycia] 427

circumductus A. Morawitz, 1862, Chlaenius 349

circumductus Audinet‑Serville, 1821 (as 1830), Dytiscus 52

circumductus Régimbart, 1899, Neptosternus 77



circumflexa Dejean, 1831, Lophyra [Cicindela] 113

circumflexus Breit, 1908, Bidessus 55

circumflexus Fabricius, 1801, Dytiscus 51



circumpicta W. Horn, 1895, Chaetodera [Cicindela] 103

circumpictula W. Horn, 1938, Chaetodera [Cicindela] 103

circumpunctatus Chaudoir, 1846, Harpalus 374

circumscripta Fischer von Waldheim, 1828, Cephalota [Cicindela] 102

circumscriptus Duftschmid, 1812, Chlaenius [Carabus] 353



circumscriptus Lacordaire, 1835, Dytiscus 51

circumseptus Germar, 1824, Calathus 524

cirenaicus A. Fiori, 1914, Paussus 210

cirocchii Magrini & Vanni, 1989, Anillus 238

cirocchii Magrini & Vanni, 1986, Duvalius 302

cirtense Netolitzky, 1914, Bembidion 252

ciscaucasicum Lutshnik, 1934, Agonum 454

ciscaucasicus Lutshnik, 1921, Harpalus 377

ciscaucasiens Ljovuschkin, 1972, Nannotrechus [Birsteiniotrechus] 315

ciseniseicus Belousov & Kabak, 1996, Trechus 335



cisteloides Panzer, 1793, Calathus [Carabus] 525

cisteloides Motschulsky, 1844, Harpalus 374

citamgineus Lassalle & Charet, 1992, Carabus 193

citharus Goulet and Smetana, 1997, Elaphrus 207



civilis Ledoux & Roux, 1998, Nebria 92

claireae W. Hansen, 1967, Catascopus 435

clairi Géhin, 1885, Carabus 140

clairi Abeille de Perrin, 1881, Duvalius [Trechus] 302

clairvillei Lapouge, 1913, Carabus 168

clairvillii Dejean, 1828, Pterostichus 503

clandestinum Sturm, 1824, Agonum 451

clandestinus Schaufuss, 1882, Carabus 163

clandestinus Schauberger, 1927, Ophonus [Harpalus] 389

clanensis Depoli, 1938, Typhlotrechus 344

clara Andrewes, 1922, Dioryche 369

clara Andrewes, 1924, Pristosia [Calathus] 531

clarescens Bates, 1892, Brachinus 216

claricollis Cavazzuti & Ratti, 1998, Carabus 128



clarissimus Roubal, 1930, Acupalpus 398

clarkii Dawson, 1849, Bembidion [Lopha] 248

clarkii Wollaston, 1862, Nebrioporus [Hydroporus] 66

clarum Andrewes, 1923, Bembidion 260



clathrata Solsky, 1871, Tenomerga [Cupes] 25

clathratum Kolenati, 1845, Calosoma 119

clathratus Linné, 1761, Carabus 160



clathratus Schaufuss, 1869, Percus 478

claudejeannei Marggi, nom. nov., Bembidion 19, 245

claudiae Deuve, 1996, Trechus 328

claudinae Laneyrie, 1967, Trichaphaenops 344

clavatus Latreille, 1804, Agabus [Dytiscus] 37

clavicornis Sharp, 1882, Agabus 37



clavicornis Motschulsky, 1856, Neocollyris [Collyris] 117

clavicornis DeGeer, 1774, Noterus [Dytiscus] 34



clavipes Bergsträsser, 1778, Abax [Carabus] [NO] 470

clavipes C. G. Thomson, 1859, Patrobus 285

clavipes L. G. Scriba, 1790, Pterostichus [Carabus] 509

cleopatrae Peyron, 1858, Herophydrus [Hydroporus] 71

clepsydra Sciaky & Wrase, 1997, Pterostichus 515

clermonti Puel, 1919, Bembidion 255

clermonti Puel, 1937, Dyschirius 224

clermonti Puel, 1923, Pogonus 286

clermonti Jeannel, 1949, Zuphium 574

clientulus Schweiger, 1969, Carabus 193

clientulus Schweiger, 1983, Ophonus 390

clivinoides Schmidt‑Göbel, 1846, Clivina [Eupalamus] 219

clops Andrewes, 1935, Bembidion 261

closanensis Jeannel, 1928, Duvalius 299

cluniensis Born, 1903, Carabus 138

clypealis C. G. Thomson, 1867, Agabus [Gaurodytes] 35



clypealis Sharp, 1876, Hydrovatus 70

clypealis Reitter, 1900, Ophonus 390

clypeata Fischer von Waldheim, 1821, Cicindela 104



clypeatus Fischer von Waldheim, 1844, Acinopus 368

clypeatus M. F. Adams, 1817, Carabus 186

clypeatus P. Rossi, 1790, Dixus [Scarites] 365

clypeatus Putzeys, 1866, Dyschirius 225

clypeatus Fischer von Waldheim, 1829, Ophonus 392

clypeonitens J. Müller, 1943, Apotomus 237



cnemerythrum Stephens, 1839, Bembidion [Peryphus] 246

cnemidotum Bates, 1883, Bembidion 254

cnesipus Andrewes, 1937, Himalosphodrus [Sphodropsis] 532

coadunata Dejean, 1825, Cymindis 412



coadunatus DeGeer, 1774, Cychrus [Carabus] 203

coarctatum C. R. Sahlberg, 1827, Bembidion 244

coarctatus LaFerté‑Sénectère, 1851, Callistomimus [Callistus] 348

coarctatus Brullé, 1836, Carabus 176

coarctatus Lucas, 1842, Poecilus 481



coarctatus J. R. Sahlberg, 1903, Tachyura [Tachys] 279

coarcticolis Reiche, 1862, Deronectes [Hydroporus] 58

coarcticollis Apfelbeck, 1904, Pterostichus 516

cobosi Mateu, 1965, Cymindis 412

cobosi Mateu, 1952, Trymosternus 410

coccinelloides L. Petagna, 1819, Omophron [Nitidula] 208

codinae Lapouge, 1913, Carabus 141

codonotus Andrewes, 1933, Dromius 420

coelestinum Motschulsky, 1844, Bembidion [Peryphus] 265

coelestinus Chaudoir, 1856, Chlaenius 356

coelestinus Motschulsky, 1865, Dicranoncus [Laxocrepis] 458

coelestinus Motschulsky, 1850, Ophonus 392

coelestis Baliani, 1932, Amara 554

coelestis Steuart, 1855, Carabus 143

coelestis Bates, 1888, Lebia 427

coelestis Sciaky & Pavesi, 1994, Trechus 328

coelesyriae Netolitzky, 1921, Bembidion 270

coelopterus Chaudoir, 1879, Metacolpodes [Colpodes] 460

coerulea Pallas, 1773, Cicindela 104



coerulea Herbst, 1806, Cylindera [Cicindela] 109

coeruleatus Fairmaire, 1891, Andrewesius [Colpodes] 456



coeruleipennis Ménétriés, 1832, Ophonus [Harpalus] 389

coeruleocephala Chaudoir, 1871, Lebia 427



coeruleocephalus Motschulsky, 1844, Chlaenius 350

coeruleolabris Mandl, 1975, Jansenia [Cicindela] 112



coeruleomicans Lebis, 1949, Carabus [Chrysocarabus] [HN] 138

coeruleomicans Sirguey, 1931, Carabus 138

coeruleovirens Sturm, 1824, Poecilus [Platysma] 484

coerulescens Motschulsky, 1844, Agonum 452

coerulescens Schilsky, 1888, Anisodactylus 361

coerulescens Motschulsky, 1844, Brachinus 213

coerulescens Schilsky, 1888, Cicindela 103

coerulescens Schilsky, 1888, Harpalus 371

coerulescens Depoli, 1929, Notiophilus 96

coeruleum Audinet‑Serville, 1821, Bembidion 245



coeruleus Schilsky, 1888, Anisodactylus 361

coeruleus Steven, 1809, Chlaenius [Carabus] 355



coeruleus Schilsky, 1888, Harpalus 376

coeruleus Dejean, 1828, Laemostenus [Pristonychus] 536



coeruleus Latreille, 1806, Leistus 84

coeruleus Brullé, 1832, Pachycarus [Ditomus] 367

cognata Chaudoir, 1846, Amara [Zabrus] 565

cognata Putzeys, 1866, Amara [Curtonotus] 558

cognata Péringuey, 1896, Clivina 220

cognata Wiedemann, 1823, Cylindera [Cicindela] 110



cognatum A. Morawitz, 1862, Bembidion 262

cognatus Wollaston, 1864, Calathus 526

cognatus Sharp, 1882, Cybister 50

cognatus Gyllenhal, 1827, Dicheirotrichus [Harpalus] 403

cognatus J. Frivaldszky, 1879, Duvalius [Anophthalmus] 299

cognatus Sharp, 1882, Laccophilus 76

cognatus Dejean, 1831, Pterostichus [Feronia] 511



cognatus Stephens, 1835, Pterostichus [Steropus] [HN] 493

cognatus Uéno, 1988, Trechiama 322



cohumilis Bernau, 1913, Carabus 184

coiffaiti Jeanne, 1981, Amara [Celia] 554

coiffaiti Jeannel, 1955, Anillidius [Nannotrechus] 290

coiffaiti Mateu, 1980, Apristus 431



coiffaiti De Wailly, 1954, Bembidion [Peryphus] 20, 269

coiffaiti Jeanne, 1972, Bembidion [Ocydromus] 256

coiffaiti Decou, 1967, Duvalius 299

coiffaiti Cabidoche, 1960, Geotrechus 310

coiffaiti A. Gaudin, 1945, Hydraphaenops 311

coiffaiti Perrault, 1990, Leistus 80

coiffaiti Mateu, 1956, Microlestes 422

coiffaiti Ledoux, 1983, Nebria 92

coiffaiti Jeanne, 1996, Platyderus 522

coiffaiti A. Gaudin, 1947, Pterostichus [Lianoe] 498

coiffaiti Jeannel, 1957, Reicheia 222

coiffaiti Jeanne, 1970, Zabrus 569

coiffaitianus A. Gaudin, 1947, Aphoenops 295

coiffaitianus Deuve, 1990, Carabus 148



colasellus Deuve, 1990, Carabus 122

colasi Paulian & Villiers, 1939, Amara 552



colasi Morvan, 1971, Bembidion [Metallina] 251

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