Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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morio Aubé, 1838, Hydroporus 62

morio Ménétriés, 1832, Idiomelas [Stenolophus] 404

morio Dejean, 1828, Omphreus 446

morio Gebler, 1847, Platynus [Anchomenus] 463

morio Duftschmid, 1812, Pterostichus [Carabus] 503

morio Ménétriés, 1832, Zabrus 572

morioformis W. J. MacLeay, 1877, Miscelus 437

morisii Vigna Taglianti & Casale, 1973, Duvalius 304

moritai Uéno, 1979, Kurasawatrechus 313

moritai Uéno, 1985, Trechiama 323

moriturum Semenov, 1900, Broscosoma 235

morneri Falkenström, 1937, Rhantus 47

moroderi Breuning, 1926, Carabus [Mesocarabus] 167

moroderi Báguena Corella, 1935, Eretes 53

morosus Lapouge, 1919, Carabus 192

morozovi Dolzhanski & Ljovuschkin, 1990, Cimmerites [Birsteiniotrechus] 297

morphocaraboides Deuve, 1989, Carabus 128

morsum Netolitzky, 1930, Bembidion 255

morula K. Daniel & J. Daniel, 1891, Nebria 93

morulus Reitter, 1884, Acupalpus 399

morvanellus Deuve, 1998, Carabus 175

morvanellus Deuve, 1996, Trechus 336



morvani Lassalle & Remond, 1998, Carabus 161

morvani Morvan, 1976, Carabus 192

morvani Deuve, 1998, Cychrus 205

morvani Zamotajlov, 1990, Deltomerus 283

morvani Casale, 1988, Laemostenus 538

morvani Ledoux & Roux, 1999, Nebria 91

morvani Lassalle, 1997, Negreum 460

morvani Deuve & Lassalle, 1983, Trigonotoma 520

morvanianum Müller‑Motzfeld, 1986, Bembidion 259

morvanianus Deuve & Quéinnec, 1985, Trechus 336

moschatum Peyron, 1858, Bembidion 268

moses Schatzmayr, 1936, Bembidion 253

moskvich Farka, 1999, Leistus 81



mosorensis J. Müller, 1930, Carabus 151

mosorensis Winkler, 1939, Cychrus 204

mosorensis J. Müller, 1930, Omphreus 445

mosorensis Jedlika, 1924, Poecilus [Pterostichus] 482

mostarensis Apfelbeck, 1904, Molops 474

motoensis Tan, 1981, Therates 115

motschoulskii Fischer von Waldheim, 1842, Calosoma [Callisthenes] 118

motschoulskyanus Schauberger, 1928, Harpalus 378

motschulskii Tschitschérine, 1896, Poecilus [Feronia] 480

motschulskyi Reitter, 1891, Agonum 451

motschulskyi Heyden, 1880, Amara 549

motschulskyi F. J. Schmidt, 1860, Anophthalmus 292

motschulskyi Csiki, 1928, Bembidion 249

motschulskyi Hrdlika, nom. nov., Brachinus 19, 216



motschulskyi Kolenati, 1845, Carabus 130

motschulskyi Andrewes, 1928, Chlaenius 351



motschulskyi Gemminger & Harold, 1868, Harpalus 377

motschulskyi Chaudoir, 1846, Nebria 86



motschulskyi Schaufuss, 1865, Pseudotaphoxenus 542

motschulskyi Iablokoff‑Khnzorian, 1978, Syntomus 433



motschulskyii Chaudoir, 1855, Scarites 234

motschuskianus Lutshnik, 1912, Poecilus [Platysma] 484

motzfeldi Belousov & Sokolov, 1994, Bembidion 246



moupinensis Breuning, 1967, Carabus 183

mourguesi Schaefer, 1969, Carabus [Autocarabus] 195

mouriesi Genest, 1983, Hydraphaenops 311

mourzinei Deuve, 1998, Amerizus 242

mourzinellus Deuve, 1998, Trechus 336



moutandoni Lapouge, 1930, Calosoma [Campalita] 120

moutandoni Buysson, 1882, Carabus 185

mouthiezi Lassalle, 1991, Carabus 153

mouthiezianoides Deuve & Kaláb, 1992, Carabus 150

mouthiezianus Deuve, 1991, Carabus 134

mouzaiensis Jeannel, 1922, Trechus [to peyerimhoffi] 326

moxiensis Deuve, 1992, Carabus 176

mtiula Zaitzev, 1927, Hydroporus 63

mtsaranus Kurnakov, 1972, Carabus 132



mucdenensis G. Hauser, 1921, Carabus [Coptolabrus] 144

muchei Jedlika, 1961, Agonum 455



muchei Jedlika, 1961, Amara [Harpalus] 562

muchei Jedlika, 1962, Amara 560

muchei Jedlika, 1961, Bembidion 244

muchei Jedlika, 1965, Bembidion [HN] 244

muchei Jedlika, 1967, Brachinus 216

muchei Jedlika, 1961, Calathus 525

muchei Breuning, 1961, Carabus 153

muchei Jedlika, 1967, Demetrias 418


muchei Jedlika, 1961, Poecilus [Pterostichus] 479

muchei Jedlika, 1961, Taphoxenus 544



mucida Jedlika, 1957, Amara 559

mucida Chevrolat, 1844, Tenomerga [Cupes] 25

muciulus Huang, 1992, Harpalus 385

mucronata Ledoux & Roux, 1999, Nebria 89

mucronatus Falkenström, 1936, Agabus [Gaurodytes] 36

mucronatus Stephens, 1828, Haliplus 32

mucronatus Jeannel, 1942, Percus 478

mucronatus Straneo, 1955, Pterostichus 498



muehlfeldi Géhin, 1885, Carabus 196

muehlfeldii Duftschmid, 1812, Pterostichus [Carabus] 488

muelleri Herbst, 1784, Agonum [Carabus] 451

muelleri Jeannel, 1926, Anophthalmus 290

muelleri A. Zimmermann, 1927, Bidessus 55

muelleri Ganglbauer, 1900, Caecoparvus [Scotodipnus] 239



muelleri Haury, 1878, Carabus 165

muelleri Magistretti, 1966, Cylindera [Cicindela] 109

muelleri Winkler, 1933, Duvalius 304

muelleri Wagner, 1929, Dyschirius 229

muelleri Ganglbauer, 1903, Laemostenus 534



muelleri Cadamuro Morgante, 1958, Paradromius [Dromius] 425

muelleri Straneo, 1936, Pterostichus [Platysma] 498



muellerianus Born, 1902, Carabus 166

muellerianus Schauberger, 1933, Ophonus [Harpalus] 389

muellerianus Schatzmayr, 1907, Orotrechus [Trechus] 317

muellneri Schmid, 1972, Arctaphaenops 296

muelverstedti Reitter, 1896, Carabus 168

muetingi Morvan, 1980, Molops 473

mugecuo Deuve, 1994, Cychrus 205



mugeko Deuve, 1992, Carabus 146

mugellii Magrini, 1982, Duvalius 304



mugeti Jedlika, 1937, Bembidion 268

mugurensis Gottwald, 1993, Carabus 149

muharnicensis Knirsch, 1927, Neotrechus 315

mujahedeeni Savich, 1999, Pterostichus 486

mukaibarai Uéno, 1958, Rakantrechus 319

mukdenensis Breit, 1933, Pterostichus 511

mukdense Kirschenhofer, 1984, Bembidion 247

mukusensis Hieke, 1997, Amara 566

mulensis Sciaky, 1997, Pterostichus 514

mulianus Deuve, 1995, Carabus 188

muliensis Deuve, 1995, Cychrus 205

mullerellus Beheim & Breuning, 1943, Carabus [Trachycarabus] 192



mulleri Reitter, 1912, Haliplus 30

mulsanti Géhin, 1885, Carabus 168

mulsanti Marseul, 1866, Harpalus 380

mulsanti Peyron, 1858, Hygrotus [Hydroporus] 72

mulsantianus A. Morawitz, 1886, Carabus 188



multicostatus Antoine, 1957, Laemostenus [Pristonychus] 535

multicostis Reitter, 1888, Carabus [Megadontus] 164

multiguttata Dejean, 1825, Lophyra [Cicindela] 113



multiguttata Fleutiaux, 1890, Lophyra [Cicindela] 113

multiguttatum Chaudoir, 1850, Omophron 208



multiguttatus Olivier, 1790, Graphipterus [Carabus] 359

multiguttatus Régimbart, 1878, Hydroporus 63



multijuncta Pic, 1925, Eurynebria 79

multilineatus Falkenström, 1922, Stictotarsus [Potamodytes] 69

multipunctata Linné, 1758, Blethisa [Carabus] 206

multipunctata Uéno, 1989, Epaphiopsis 309

multipunctatum Motschulsky, 1850, Bembidion [Peryphus] 255

multipunctatus Kabak, 1997, Amara [Curtonotus] 560



multipunctatus Kraatz, 1890, Carabus 160

multipunctatus Wehncke, 1875, Haliplus 30

multipunctatus Dejean, 1828, Pterostichus [Feronia] 503



multiseriatus G. Hauser, 1921, Carabus [Damaster] 148

multiseriatus G. Hauser, 1921, Carabus [Damaster] 148

multiseta Straneo, 1984, Pterostichus [Amolopsa] 494



multisetosus C. G. Thomson, 1884, Harpalus 380

multistriatus G. Hauser, 1913, Carabus [Apotomopterus] [HN] 126

multistriatus Lapouge, 1925, Carabus 170

multistriatus Motschulsky, 1866, Carabus 176

multistriolatus Guignot, 1956, Copelatus 48

multisulcatum Reitter, 1890, Bembidion 268



multominor Heer, 1838, Pterostichus 504

muluyensis Antoine, 1957, Orthomus 477

mumbugianus Mateu, 1985, Trechus 336

munakatai Ishikawa, 1968, Carabus [Damaster] 122

munakataorum Ishikawa, 1969, Carabus [Procrustes] 164



munda Holdhaus, 1942, Amara 563

mundatum Netolitzky, 1920, Bembidion 247

mundatus Jedlika, 1940, Pterostichus 513

mundus Germar, 1836, Platynus [Agonum] 463

mundus Jedlika, 1938, Pterostichus 501

munelensis Apfelbeck, 1906, Molops 474

munganasti Reitter, 1908, Bembidion 255

munganasti Reitter, 1905, Carabus 146



munganasti Reitter, 1908, Harpalus 381

munganasti Reitter, 1905, Leistus 83

municipalis Duftschmid, 1812, Amara [Carabus] 557



munsteri Hellén, 1935, Agonum [Europhilus] 453

munsteri Larsson & Gígja, 1959, Nebria 89



munsterii Helliesen, 1890, Hydroporus 62

munzurensis Cavazzuti & Lassalle, 1987, Carabus 185

murakamii Uéno, 1997, Ishikawatrechus 312

murakamii Uéno, 1984, Trechiama 323

murakii Ishikawa & Kubota, 1984, Carabus 177

muralevitshi Lutshnik, 1915, Poecilus [Platysma] 482

muranishii Tanaka, 1958, Pterostichus 493

murex Coquerel, 1859, Poecilus [Feronia] 483

murgabica Tschitschérine, 1902, Amara 566

murgabicus Kryzhanovskij & Mikhailov, 1987, Dromius 420

muriauxi Jeannel, 1957, Duvalius 304



murinus Sturm, 1835, Deronectes [Hydroporus] 58

murinus Régimbart, 1892, Orectochilus 28

muro Ishikawa & Kubota, 1995, Carabus 178

murreense Müller‑Motzfeld, 1985, Bembidion 257

murzini Zamotajlov, 1999, Deltomerodes 281

murzini Farka, 1999, Leistus 81

murzini Ledoux & Roux, 1999, Nebria [add] 90

murzorum Belousov & Kabak, 1994, Trechus 336

mus Jedlika, 1965, Skoeda [Agonum] 466

musartianus Gottwald, 1987, Carabus 145

musashiensis Kasahara, 1993, Pterostichus 502

musayabensis Ali, 1967, Ophonus 391

muscorum P. Moravec & Wrase, 1998, Trechus 336

musculum Jedlika, 1965, Xestagonum [Agonum] 468

mushai Jedlika, 1951, Lebia 429

musicus Klug, 1834, Herophydrus [Hydroporus] 71



mussardi Antoine, 1951, Carabus [Hygrocarabus] 146

mussardi Saha & Sengupta, 1979, Chlaenius 354

mussardi Antoine, 1962, Geocharis [DA] 239



mussini Germar, 1824, Carabus 200

mustella Andrewes, 1923, Clivina 220



mutabilis Géhin, 1885, Carabus 194

mutabilis Reitter, 1896, Carabus [HN] 131

mutabilis Reiche, 1855, Syntomus [Metabletus] 432

mutator Bates, 1883, Negreum [Colpodes] 460



mutator Poppius, 1906, Pterostichus [Cryobius] 491

mutatus Gemminger & Harold, 1868, Herophydrus [Hydroporus] [add, syn of guineensis] 71

mutatus Bedel, 1876, Trechus 326

mutilatus Fabricius, 1801, Aptinus [Brachinus] 212

mutilatus Rosenhauer, 1856, Pseudotrechus 438

mutinensis Branden, 1885, Dytiscus 52

mutinensis Pederzani, 1971, Dytiscus 52


mutsuomiyatakei Habu, 1974, Hikosanoagonum [Platynus] 458

myohyangensis Deuve & Li, 1998, Carabus 122

myohyangicus Deuve & Li, 1999, Carabus 174

myops Peyerimhoff, 1929, Mesolestes [Dromius] 421

myrmidon Fairmaire, 1859, Metadromius [Dromius] 421

myrrha J. Thomson, 1857, Cylindera [Cicindela] 112

mysorensis Guignot, 1954, Neohydrocoptus [Hydrocoptus] 34

mystica Casey, 1918, Amara 549

nacharensis Rost, 1893, Carabus [Plectes] 200



nactum Casey, 1918, Bembidion 263

nadari Vuillefroy, 1893, Pterostichus [Feronia] 498

naeviger A. Morawitz, 1862, Chlaenius 353

naevius Gmelin, 1790, Agabus [Dytiscus] 40

naevulus Reitter, 1919, Brachinus 214

nagahinis Uéno, 1976, Trechiama 323

naganoensis Tanaka, 1958, Pterostichus 502

nagpurensis Bates, 1891, Dioryche [Platymetopus] 370

nagpurensis Bates, 1891, Loxoncus [Anoplogenius] 404

nagyagensis Birthler, 1886, Carabus 151

naiman Belousov & Kabak, 1992, Trechus 341

nainensis Casey, 1918, Amara [Bradytus] 554

naini Jedlika, 1964, Microlestes [Metabletus] 423



nairica Iablokoff‑Khnzorian, 1964, Amara 561

nairicum Iablokoff‑Khnzorian, 1976, Bembidion 255

nairicus Pavesi & Sciaky, 1992, Trechus 336

nakabusei Jedlika, 1958, Bembidion 268

nakaguroi Uéno, 1960, Trechus 336

nakamurai Ishikawa, 1966, Carabus [Apotomopterus] 177



nakamurai Ishikawa, 1968, Carabus 149

nakamurai V. B. Guéorguiev, 1970, Copelatus 48

nakanei Jedlika, 1965, Bembidion 265

nakanei Nilsson, 1994, Ilybius 42

nakanei Kasahara, 1986, Onycholabis 461

nakanei Nilsson, 1997, Platambus [Agabus] 44

nakanei Straneo, 1955, Pterostichus 502

nakanoshimense Uéno, 1955, Bembidion 243

nakaoi Uéno, 1972, Trechiama 323

nakatomii Ishikawa, 1966, Carabus 159

nakayamai Morita, 1992, Badister 441

nakeralae Reitter, 1883, Troglocimmerites [Anophthalmus] 344



nakrensis Novotný & Voíšek, 1988, Carabus 200

naldii Ghidini, 1932, Trechus 336



nalinie Vazirani, 1969, Hyphydrus 74

naloni Salgado Costas, 1993, Apoduvalius 295

naltarense Müller‑Motzfeld, 1988, Bembidion 261


namanganensis Heyden, 1886, Carabus 147

namanganensis Jedlika, 1946, Cymindis 416

namanganensis Heyden, 1885, Microderes [Pangus] 387

namanganensis Kabak, 1994, Poecilus 484

nambinensis Boldori, 1935, Duvalius 304

namchabarwana Imura, 1999, Cychropsis 202

namhaedoensis Kwon & Lee, 1984, Carabus 183

namigatai Uéno, 1989, Trechiama 323

namrum Jedlika, 1963, Pterostichus 491

namrun Jedlika, 1963, Platyderus 523

namunlaensis J. Schmidt, 1998, Trechus 332

nana Ledoux & Roux, 1996, Nebria 91

nana Bates, 1889, Perigona 448

nana Gyllenhal, 1810, Tachyta [Bembidium] 277

nangnimicus Blumenthal & Deuve, 1984, Carabus 134

nangqenicus Deuve, 1995, Carabus 150

nanhutanus Habu, 1973, Apatrobus [Patrobus] 280

nankotaizanus Kanô, 1932, Carabus 143

nanlingensis Deuve & Tian, 1999, Carabus 127

nanlingensis Deuve & Tian, 1999, Leistus 81



nanniscus Péringuey, 1896, Laemostenus [Pristonychus] 536

nanniscus Péringuey, 1896, Polyderis [Tachys] 274

nannus Jeannel, 1931, Duvalius 299

nanophyes Andrewes, 1925, Elaphropus [Tachys] 273

nanosomus G. Hauser, 1931, Carabus [Apotomopterus] 128

nanping Farka, 1999, Leistus 81

nanpingense Kirschenhofer, 1984, Bembidion 244

nanpingensis Lassalle, 1999, Acalathus [Procalathus] 529

nanpingensis Deuve, 1992, Carabus 128

nanpingensis Kirschenhofer, 1999, Chlaenius 354

nanpingensis Toledo & Mazzoldi, 1996, Hydroporus 63

nanpingensis Zamotajlov & Sciaky, 1996, Parapenetretus 284

nanpingensis Kirschenhofer, 1997, Synuchus 545

nanpingicus Deuve, 1995, Carabus 121

nanschanicus Semenov, 1898, Carabus 175

nanshanica Shilenkov, 1982, Nebria 91

nantouensis Kirschenhofer, 1996, Parena 434

nanulus Tschitschérine, 1898, Microderes [Harpalus] 387

nanum Jedlika, 1965, Agonum 455

nanum Stephens, 1835, Bembidion [Lopha] 249

nanus Gmelin, 1790, Bidessus [Dytiscus] 55

nanus Kraatz, 1897, Carabus [Deuterocarabus] 130

nanus Putzeys, 1866, Dyschirius 227

nanus A. Fiori, 1914, Metadromius [Dromius] 421

nanus Mateu, 1953, Microlestes 423


nanus Belousov, 1998, Paracimmerites 318

nanus Heer, 1837, Pterostichus [Argutor] 492

nanus Habu, 1954, Trichotichnus 395

nanwutai Kleinfeld, Korell & Wrase, 1996, Carabus 190

nanxiangensis Deuve & Tian, 1999, Carabus 127

napaea Kasahara, 1988, Pterostichus 502

napoleonius Reiche, 1857, Patrobus 285

napraensis Belousov & Zamotajlov, 1993, Carabus 132

narae Lindroth, 1956, Synuchus 545

naraensis Uéno, 1955, Perileptus 289

naraensis Uéno, 1953, Polyderis 274

naraglavi Mlejnek & J. Moravec, 1995, Anophthalmus 22, 292

naranensis Heinertz, 1978, Carabus 155

naratensis Deuve, 1993, Carabus 178

naratensis Deuve, 1993, Trechus 336

naraticus Deuve, 1993, Carabus 123

narense J. Schmidt, 1998, Xestagonum 467

narentinus A. Zimmermann, 1915, Graptodytes [Hydroporus] 59

narentinus Reitter, 1884, Lymnastis 274

narentinus Apfelbeck, 1904, Molops 473

narentinus Drovenik & Peks, 1999, Stenolophus 23, 406

narosnyi Csiki, 1946, Carabus 185

narukawai Morita, 1989, Apatrobus 280

narukawai Ishikawa & Kubota, 1995, Carabus 178

narukawai Morita, 1997, Trichotichnus 396

narynensis Csiki, 1927, Carabus 145

narynensis Jedlika, 1958, Harpalus 377

narynensis Kabak, 1997, Poecilus 484

narynensis Belousov & Kabak, 1992, Trechus 336

narynicus Obydov, 1999, Carabus 182

narzikulovi Kryzhanovskij, 1972, Bembidion 257

narzikulovi Mikhailov, 1972, Parazuphium 574



nastasi Pavievi, 1995, Carabus 152

nasui Kasahara, 1993, Pterostichus 513

nasutus Sharp, 1887, Bidessus 55

natalensis Putzeys, 1861, Clivina 220

natalensis Chaudoir, 1871, Lebia 429



natalensis Péringuey, 1896, Selina [Ega] 407

natalia Péringuey, 1898, Drypta 358

nataliae Kryzhanovskij, 1974, Amara [Curtonotus] 560

nataliae Kabak & Putchkov, 1996, Nebria 91

natator Linné, 1758, Gyrinus 27

natrix Sharp, 1884, Oreodytes [Hydroporus] 68

natruc Farka, 1998, Leistus 81



natvigi Csiki, 1929, Amara 550

nauceras Benson, 1846, Paussus 210

naumanni Breuning, 1971, Carabus 131

nauzieli Fairmaire, 1859, Hydaticus 53

navarica Zaballos & Wrase, 1998, Typhlocharis 241

navaricus Jeanne, 1985, Brachinus 213

navaricus Coiffait & A. Gaudin, 1950, Hydraphaenops [Aphoenops] 311

navaricus Coiffait, 1958, Hypotyphlus [Microtyphlus] 239

navaricus Vuillefroy, 1893, Laemostenus [Antisphodrus] 535



navaricus Dejean, 1828, Percus [Feronia] 479

navaricus Colas, 1951, Pterostichus [Haptoderus] 489

navaricus Jeannel, 1937, Pterostichus [Haptoderus] 489

navaricus Vuillefroy, 1869, Trechus [Anophthalmus] 336



navarricus Lapouge, 1925, Carabus 194

navasi Vidal y López, 1918, Cicindela 106

navatensis Novoa, 1975, Metadromius 422

naviauxi Morvan, 1999, Aparupa 456

naviauxi Wrase, 1995, Badister 440

naviauxi Jeanne, 1972, Bembidion [Nepha] 253

naviauxi Deuve & Mourzine, 1998, Cychrus 205

naviauxi Morvan, 1995, Skoeda 466

naviauxi Morvan, 1995, Xestagonum [Colpodes] 468

naxiana Apfelbeck, 1904, Cymindis 414

naxius J. Müller, 1924, Gyrinus 28

nazarovi Lafer, 1976, Platynus [Agonum] 463



nazmovi Obydov, 1999, Carabus 165

neapolitanus Leoni, 1907, Laemostenus 532

neapolitanus Reiche, 1855, Platyderus [Feronia] 523


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