Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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nihilumalbi Schmid, 1972, Arctaphaenops 295

nihozana Bates, 1883, Cicindela 107

niigatanus Schauberger, 1929, Harpalus 385

niitakaensis Minowa, 1932, Leistus 83



niitakana Kanô, 1931, Cylindera [Cicindela] 108

niitakana Kanô, 1930, Nebria 92



nikkoense Bates, 1883, Bembidion 271

nikkoensis Bates, 1883, Trephionus 547

nikodymi Dvoák, 1995, Pterostichus 505

nikolajevi Kabak, 1998, Carabus 146

nikolajevi Kabak, l997, Cymindis 418

nikolajevi Belousov & Kabak, 1992, Trechus 336

nikolajewi Hieke, 1999, Amara 557

nikolskyi Jedlika, 1932, Ophonus 391


nila Andrewes, 1924, Amara 550

nilghiricus Régimbart, 1903, Hyphoporus 73



niliusi Breuning, 1927, Carabus 198

nilotica Dejean, 1825, Habrodera [Cicindela] 112

nilotica Chaudoir, 1871, Lebia 429

niloticum Dejean, 1831, Bembidion 253



niloticus Waltl, 1838, Gyrinus 27

niloticus Motschulsky, 1862, Lymnastis 274

nilssoni Fery & Wewalka, 1992, Deronectes 58

nilssoni Wewalka, 1997, Microdytes 75

nimbatidius Chaudoir, 1878, Pterostichus [Feronia] 513

nimbatus A. Morawitz, 1862, Pterostichus 513

ninetopa Nakajima, 1993, Carabus [Damaster] 122

ninini Marion, 1989, Lionychus 432

niohozana Bates, 1883, Nebria 94

niponensis Bates, 1883, Calomera [Cicindela] 99

niponensis Bates, 1873, Clivina 220

niponensis Brinck, 1941, Gyrinus 27

niponensis Kamiya, 1939, Japanolaccophilus [Neptosternus] 76

niponensis Bates, 1883, Nipponoharpalus [Harpalus] 388

niponensis Bates, 1883, Panagaeus 447

niponensis Lewis, 1888, Yamatosa [Rhysodes] 78

niponicus Csiki, 1927, Carabus 148

niponicus Lewis, 1879, Notiophilus 97

nipponensis Jaeger & Wrase, 1994, Bradycellus 402



nipponensis Mandl, 1992, Chlaenius 351

nipponensis Habu, 1964, Parena 434



nipponica Habu, 1959, Amara 567

nipponica Habu, 1963, Archicolliuris 443

nipponica Habu, 1950, Jujiroa [Sphodropsis] 459

nipponicum Habu, 1972, Agonum 453

nipponicum Habu & Uéno, 1955, Bembidion [Hydrium] 243

nipponicus Habu, 1983, Dromius 420

nipponicus Habu & Baba, 1967, Elaphropus 273

nipponicus Habu, 1950, Ishikawatrechus 312

nipponicus Takizawa, 1933, Nebrioporus [Deronectes] 66

nipponicus Habu, 1955, Synuchus [Calathus] 545

nipponicus Habu, 1961, Trichotichnus 396



nirensis Heinz, 1970, Carabus 157

nishiawakurae Habu, 1980, Apatrobus [Patrobus] 280

nishidai Deuve & Imura, 1993, Carabus 135

nishidai Nakane, 1989, Pterostichus 515

nishijimai Imura, 1991, Carabus 122

nishikawai Ishikawa, 1966, Carabus [Apotomopterus] 177

nishikawai Uéno, 1987, Epaphiopsis 309


nishikawai Uéno, 1980, Himalaphaenops 311

nishikawai Uéno & Saito, 1991, Jujiroa 459

nishikawai Uéno, 1993, Kurasawatrechus 313

nishikawai Uéno, 1998, Libotrechus 314

nishikawai Uéno, 1980, Stygiotrechus 320

nishikawai Uéno, 1991, Trechus 325

nishioi Habu, 1961, Trichotichnus 396

nishiyamai Kasahara, 1986, Pterostichus 502



nitens Putzeys, 1866, Amara [Curtonotus] 560

nitens Gautier des Cottes, 1866, Carabus 187

nitens Linné, 1758, Carabus 153



nitens Faldermann, 1836, Chlaenius 350

nitens Andrewes, 1935, Cymindis 414



nitens LeConte, 1858, Dicheirotrichus [Bradycellus] 403

nitens Falkenström, 1939, Hydroglyphus [Bidessus] 56

nitens Jacquemet, 1882, Lophyra [Cicindela] 113

nitens Chaudoir, 1850, Poecilus [Feronia] 479

nitens Andrewes, 1929, Scarites 233

nitens Motschulsky, 1864, Stenolophus [Egadroma] 405

nitens Heer, 1837, Trichotichnus [Harpalus] 396

nitida Sturm, 1825, Amara 550

nitida Lichtenstein, 1796, Cicindela 104

nitida Mjöberg, 1905, Clivina 220



nitida Wiedemann, 1821, Cylindera [Cicindela] 111

nitidicollis Dejean, 1826, Chlaenius 351

nitidicollis Harald Lindberg, 1953, Gomerina [Platyderus] 521

nitidicollis Motschulsky, 1844, Poecilus 483

nitidicollis Sciaky & Pavesi, 1997, Pseudotaphoxenus 541

nitididorsus Ishikawa & Kim, 1983, Carabus 183



nitidior Reitter, 1881, Carabus 187

nitidior Pic, 1924, Licinus 441

nitidipunctatus Ishikawa, 1966, Carabus [Procrustes] 164

nitidiuscula Putzeys, 1873, Amara [Celia] 556

nitidula Fabricius, 1787, Nebria [Carabus] 89

nitidula A. Morawitz, 1862, Pristosia [Calathus] 531



nitidulum Marsham, 1802, Bembidion [Carabus] 259

nitidulus Mulsant & Wachanru, 1852, Brachinus 214

nitidulus Sharp, 1882, Canthydrus 34



nitidulus Laporte, 1834, Catascopus 435

nitidulus Schrank, 1781, Chlaenius [Carabus] 350



nitidulus Thunberg, 1784, Chlaenius [Carabus] 350

nitidulus Dejean, 1825, Dixus [Ditomus] 365

nitidulus Motschulsky, 1844, Harpalus 372

nitidulus Dejean, 1826, Hololeius [Chlaenius] 356

nitidulus G. R Waterhouse, 1833, Notiophilus 97

nitidulus Normand, 1916, Pseudanillus [Anillus] 238

nitidulus Motschulsky, 1864, Stenolophus 406

nitidulus Piochard de la Brûlerie, 1868, Syntomus [Metabletus] 433

nitidum Motschulsky, 1844, Agonum 451

nitidus Faldermann, 1836, Acinopus 368

nitidus Stephens, 1835, Acupalpus [Trechus] 398

nitidus Fabricius, 1801, Agabus [Dytiscus] 38

nitidus Mjöberg, 1905, Distichus [Clivina] 231

nitidus Motschulsky, 1864, Dregus 370

nitidus Dejean, 1825, Dyschirius [Clivina] 226



nitidus Sturm, 1818, Harpalus 377

nitidus Uéno, 1956, Ishidatrechus 312



nitidus Sturm, 1835, Laccornis [Hydroporus] 75

nitidus Motschulsky, 1865, Laemostenus [Pristonychus] [jun. HN?] 537

nitidus Schaufuss, 1865, Laemostenus [Pristonychus] [jun. HN?] 538

nitidus Duftschmid, 1812, Leistus [Carabus] 82



nitidus Motschulsky, 1848, Neophygas [Phygas] 387

nitidus G. R. Waterhouse, 1833, Notiophilus 96

nitidus Lagar Mascaró, 1965, Percus 479

nitidus Mateu, 1957, Philorhizus 426

nitidus Dejean, 1828, Poecilus [Feronia] 479

nitidus Chaudoir, 1843, Scarites 233

nitidus Motschulsky, 1861, Synuchus [Crepidactyla] 546



nitidus Motschulsky, 1845, Zabrus 572

nitouense Jedlika, 1934, Xestagonum [Colpodes] 468

nitouensis Jedlika, 1937, Pristosia [Eucalathus] 531

nivale Heer, 1837, Bembidion 20, 269



nivale Breuning, 1927, Calosoma 121

nivalis J. Müller, 1922, Anophthalmus [Trechus] 292

nivalis Heer, 1839, Hydroporus 63

nivalis Apfelbeck, 1902, Laemostenus 534

nivalis Apfelbeck, 1890, Molops [Stenochoromus] 475

nivalis Paykull, 1798, Nebria [Carabus] 18, 88

nivalis Apfelbeck, 1904, Platyderus 522

nivalis Apfelbeck, 1904, Pterostichus 499

nivalis Brisout de Barneville, 1863, Pterostichus [Argutor] 490

nivalis R. F. Sahlberg, 1844, Pterostichus [Feronia] 491



nivalis Panzer, 1796, Synuchus [Carabus] 546

nivalis Uéno, 1986, Trechiama 323



niveus Géhin, 1885, Carabus 166

nivicinctoides W. Horn, 1892, Callytron [Cicindela] 101

nivicinctum Chevrolat, 1845, Callytron [Cicindela] 101

nivicola Andrewes, 1923, Bembidion 266

nivicola Lapouge, 1930, Calosoma [Campalita] 121

nivicola Semenov, 1891, Cymindis 415


nivicola Ménétriés, 1832, Pterostichus [Feronia] 501

nivicola Chaudoir, 1846, Trechus 336



nivina Gistel, 1857, Amara [Celia] 556

nivium Tschitschérine, 1900, Amara 550

nivium Breuning, 1933, Carabus 150

nivium Andrewes, 1925, Leistus 81

nivium Bates, 1891, Xestagonum [Colpodes] 468

nivosus Heer, 1837, Carabus 181

nobile Rottenberg, 1870, Bembidion 249

nobilis Duftschmid, 1812, Amara [Carabus] 563

nobilis Sturm, 1825, Amara 566

nobilis Dejean, 1831, Brachinus 212

nobilis Dejean, 1828, Broscus [Cephalotes] 236

nobilis Ganglbauer, 1886, Carabus 165

nobilis Schaufuss, 1882, Carabus 163

nobilis Jedlika, 1963, Coptodera 436

nobilis Andrewes, 1933, Cymindis 414

nobilis Xie & Yu, 1991, Euschizomerus 447

nobilis Wollaston, 1864, Masoreus 357

nobilis Poppius, 1905, Oreodytes [Hydroporus] 67

nobilis Goeze, 1777, Panagaeus [Carabus] 447

nobilis Stierlin, 1869, Pterostichus 504

nobilitatus Faldermann, 1836, Harpalus 381

nobuchii Jedlika, 1964, Tachys [Trechus] 276

nobuyoae N. Ito, 1999, Trichotichnus 394

nocticolor Deuve & Kaláb, 1992, Carabus 175

noctirugulosus Deuve & Kaláb, 1996, Carabus 175



noctivaga Hochhuth, 1871, Amara 568

noctivagus Deuve, 1992, Carabus 150

noctivagus Peyerimhoff, 1927, Psammodromius 426

noctuabundus Habu, 1954, Trichotichnus 396



nocturnus Schweiger, 1969, Carabus 198

nodicollis Bates, 1874, Pronyssa 115



nodicornis Lapouge, 1908, Carabus 130

nodicornis Fairmaire & Laboulbène, 1854, Pterostichus [Feronia] 507



nodifrons Penecke, 1914, Dyschirius 226

nodulosus Creutzer, 1799, Carabus 154



noesskei Emden, 1932, Carabus 137

noesskei Bänninger, 1932, Leistus 83

noesskei Apfelbeck, 1908, Neotrechus [Trechus] 315

noesskei Apfelbeck, 1912, Neotrechus [Anophthalmus] 315

noesskei Hänel, 1940, Pterostichus 493

noguchii Bates, 1873, Chlaenius 354

noguchii Bates, 1873, Pterostichus 499

nokoensis Minowa, 1932, Leistus 81



nomax F. Balfour‑Browne, 1911, Haliplus 31

nomurai N. Ito, 1992, Anisodactylus 361

nomurai Uéno, 1960, Rakantrechus 319

nomurai Uéno, 1998, Trechus 337

nonensis Uéno, 1995, Trechiama 323

nonfriedi Reitter, 1898, Philorhizus [Dromius] 425



nonsignatus Krynicki, 1832, Harpalus [Anisodactylus] 384

nonsulcatus A. Zimmermann, 1919, Dytiscus 52

nonveilleri Scheibel, 1933, Anophthalmus 291

nonveilleri J. Müller, 1930, Trechus 337

nordafricanum De Monte, 1949, Bembidion 267



nordenskioeldii J. R. Sahlberg, 1880, Poecilus [Feronia] 481

nordmanni Chaudoir, 1844, Bembidion [Peryphus] 268

nordmanni Chaudoir, 1848, Carabus 158

nordmanni Chaudoir, 1848, Cicindela 106

nordmanni A. Morawitz, 1862, Synuchus [Taphria] 546

nordqvisti J. R. Sahlberg, 1887, Pterostichus [Feronia] 491



norensis Pham, 1977, Carabus 171

norica Schauberger, 1927, Nebria 86



noricus K. Daniel & J. Daniel, 1891, Calathus 527

noricus Sokolá, 1910, Carabus 135

noricus Ganglbauer, 1891, Pterostichus 508

noricus Meixner, 1911, Trechus 337

normandi De Monte, 1947, Bembidion 258



normandi Bedel, 1898, Cephalota [Cicindela] 102

normandi Puel, 1937, Dyschirius 227

normandi Régimbart, 1903, Hydroporus 63

normandi Jeannel, 1957, Reicheia 222

normannensis Sirguey, 1931, Carabus 139

normanneus Sirguey, 1931, Carabus 139

normannum Dejean, 1831, Bembidion 249

normodynamus Cavazzuti, 1995, Carabus 125

nortoni Andrewes, 1930, Diplous 283



norvegicus Munster, 1922, Brychius 30

norvegicus Munster, 1923, Dyschirius 226

nossibiana Fairmaire, 1895, Calleida 409

notabilangulus N. Ito, 1998, Trichotichnus 394

notabilis Roeschke, 1898, Carabus 181

notabilis Xie & Yu, 1991, Craspedophorus 447

notabilis LeConte, 1850, Hydroporus 63

notabilis Uéno, 1960, Masuzoa 314

notabilis Martínez y Saez, 1873, Zabrus 569

notarum Uéno, 1980, Stygiotrechus 320



notasicus Aubé, 1838, Cybister 51

notaticollis Aubé, 1837, Rhantus [Colymbetes] 46



notatipennis Pic, 1909, Dineutus [Dineutes] 26

notatum Andrewes, 1922, Bembidion 257

notatus Mulsant & Rey, 1861, Acupalpus 398

notatus Bergsträsser, 1778, Agabus [Dytiscus] 40

notatus Stephens, 1828, Harpalus 381

notatus Sturm, 1835, Hydroporus 63

notatus Stephens, 1827, Philorhizus [Dromius] 426

notatus Coquerel, 1859, Platyderus [Feronia] 523

notha Antoine, 1936, Amara 552

nothus M. F. Adams, 1817, Carabus 169

nothus Jeannel, 1960, Trechus 337

notmani Casey, 1924, Bembidion 266

notoi Uéno, 1981, Trechiama 323

notos Uéno, 1991, Epaphiopsis 309

notsui Uéno, 1983, Kurasawatrechus 313

notula Erichson, 1843, Canthydrus [Hydrocanthus] 34

notulatoides Xie & Yu, 1991, Dischissus 447

notulatus Fabricius, 1801, Dischissus [Carabus] 447

nouei Pic, 1932, Nebria 86

noui Jeannel, 1942, Pterostichus 498

nouristanensis Ledoux, 1985, Nebria (as noristanensis) 90

nouristanensis Casale & Ledoux, 1996, Sphodropsis 543

nouristani Ledoux, 1977, Carabus 155

novaki J. Müller, 1898, Carabus 196

novaki J. Müller, 1911, Duvalius [Trechus] 306

novaki J. Müller, 1930, Molops 473

novaki Všeteka, 1929, Nebria 86

novaki Mlejnek, J. Moravec & Udral, 1994, Orotrechus 317

novaki J. Müller, 1934, Parazuphium [Zuphium] 573

novemlineatus Stephens, 1829, Hygrotus [Hydroporus] 72



novenoei Morvan, 1976, Trechus 329

novenumus Deuve, 1995, Carabus 126

novitia Bates, 1883, Cylindera [Cicindela] 110

novorossicus Semenov, 1899, Acinopus 368

novotnyorum Vysoký, 1981, Pterostichus 496

nowitzkii Tschitschérine, 1899, Pterostichus [Feronia] 510

nox Semenov, 1897, Cylindera [Cicindela] 109

nozakicola Imura & Mizusawa, 1994, Carabus 177

nubatama Habu, 1957, Setolebia [Lebia] 430



nubicola Zolotarev, 1913, Carabus [Plectes] 132

nubicola Tschitschérine, 1903, Leistus 81



nubiensis Régimbart, 1895, Agabus 38

nubiferum A. Morawitz, 1862, Asaphidion [Tachypus] 243

nubigena Haliday, 1838, Calathus 527

nubila Andrewes, 1931, Anchista 408



nubilipennis Fairmaire, 1888, Pristosia [Calathus] 531

nubilus Régimbart, 1899, Hydaticus 53

nubilus Gmelin, 1790, Scarodytes [Dytiscus] 68

nubivagus Wollaston, 1864, Leistus 83

nuceus Fairmaire, 1887, Platynus [Anchomenus] 463

nuda W. Horn, 1905, Salpingophora [Cicindela] 114

nudicollis Peyerimhoff, 1911, Nebria 85

nudilabrus J. Müller, 1898, Carabus 196

nudopectoralis W. Horn, 1903, Myriochila [Cicindela] 113

nudoscripta W. Horn, 1915, Cylindera [Cicindela] 110

nuessleri Gebert, 1991, Cephalota 103

nugax Lompe, 1997, Trechus 337

nukatli Belousov, 1990, Trechus 333

numatai Chûjô, 1966, Tachyura [Tachys] 280

numatai Uéno, 1967, Trechus 337

numida Jeannel, 1957, Reicheia 222

numidica Lucas, 1846, Lebia 427

numidicum Lucas, 1846, Agonum [Anchomenus] 451

numidicum Lucas, 1846, Zuphium 574



numidicus Gautier des Cottes, 1866, Calathus 524

numidicus Bedel, 1925, Canthydrus 33

numidicus Putzeys, 1846, Dyschirius 228

numidicus Bedel, 1893, Harpalus 379

numidicus A. Fiori, 1913, Leistus 84

numidicus Normand, 1941, Microlestes 423



numidicus Lucas, 1842, Poecilus 480

numidicus Lucas, 1846, Porotachys [Bembidium] 274

numidicus Bedel, 1889, Rhithrodytes [Hydroporus] 68

numidus Laporte, 1834, Carabus 152

numidus Chaudoir, 1859, Orthomus [Feronia] 476

nuncaestimatum Netolitzky, 1939, Bembidion 265

nuptialis Csiki, 1912, Duvalius [Anophthalmus] 299

nurbeki Kaláb, 1997, Carabus 147



nurensis Morvan, 1981, Laemostenus [Pristonychus] 538

nuristanica Mandl, 1955, Cicindela 108

nuristanicum Fassati, 1957, Bembidion 261

nuristanicus G. Ochs, 1955, Orectochilus 28

nuristanus Jedlika, 1967, Brachinus 216

nuristanus Jedlika, 1956, Chlaenius 356

nuristanus Jedlika, 1956, Prakasha [Harpalus] 393

nuytenae Kirschenhofer, 1981, Agonum 451

nyalamensis Deuve, 1996, Carabus 176

nyingtri Farka, 1998, Leistus 81

oasis Wewalka, 1992, Hydroporus 63



obbergeni G. Hauser, 1914, Carabus [Coptolabrus] 144

obenbergeri Lutshnik, 1935, Amara 566

obenbergeri Lutshnik, 1928, Bembidion 244



obenbergeri íha, 1952, Brychius 30

obenbergeri Semenov & Znojko, 1932, Carabus 190



obenbergeri Maan, 1935, Dyschirius 224

obenbergeri Jedlika, 1928, Harpalus 371

obenbergeri Jedlika, 1963, Orionella 438

obenbergeri Jedlika, 1934, Platynus [Colpodes] 465

obenbergeri Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

obenbergeri Kult, 1940, Stomis 518

obenbergeri Maan, 1932, Tapinopterus [Pterostichus] 520

obenbergerianus Breuning, 1927, Carabus 168

oberleitneri Dejean, 1831, Percus [Feronia] 478

oberleitneri Dejean, 1830, Siagona 19, 218



oberthueri Venet, 1930, Carabus 168

oberthueri Fleutiaux, 1894, Cosmodela [Cicindela] 108

oberthueri Beuthin, 1904, Cylindera [Cicindela] 109

oberthueri Reitter, 1885, Leistus 82

oberthueri Heller, 1903, Lioptera 436

oberthueri Ledoux & Roux, 1991, Nebria 91

oberthueri Gestro, 1892, Omophron 208

oberthueri Tschitschérine, 1894, Trigonotoma 520

oberthuri Pater, 1938, Ophonus [Metophonus] 388

oberti Motschulsky, 1864, Dromius 420

obesa Baliani, 1940, Amara 560

obesicollis Putzeys, 1861, Clivina 220

obesulus LeConte, 1852, Harpalus 377

obesus Schönherr, 1806, Acinopus [Carabus] 368

obesus Colas, 1938, Calathus 528



obesus A. Morawitz, 1862, Harpalus 379

obesus Abeille de Perrin, 1886, Trichaphaenops [Anophthalmus] 344

obesus Audinet‑Serville, 1821, Zabrus [Harpalus] 569

obesus Stephens, 1828, Zabrus 573

obirensis Jeannel, 1926, Anophthalmus 291

obirensis Schauberger, 1921, Pterostichus [Lyperophorus] 508

obirensis Schönmann, 1937, Trechus 330



obligata Casey, 1918, Amara [Celia] 565

obliqua Motschulsky, 1850, Cymindis [Tarus] 412

obliquatus Fairmaire, 1881, Catascopus 435

obliquatus Uéno, 1997, Ishikawatrechus 312



obliquebasalis Jedlika, 1962, Pterostichus 505

obliquebasalis Breit, 1914, Trechus 337



obliquefasciata Fischer von Waldheim, 1828, Cicindela 104

obliquefasciata M. F. Adams, 1817, Cylindera [Cicindela] 109

obliquefasciatus W. Horn, 1912, Therates 116


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