Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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simulans J. R. Sahlberg, 1880, Amara [Harpalus] 560

simulans Bedel, 1895, Neolaphyra [Cicindela] 114

simulator Kraatz, 1876, Carabus 174

simulator J. Müller, 1922, Dyschirius 226

simulator Westhoff, 1882, Graphoderus [Hydaticus] 49

simulator Bänninger, 1933, Nebria 93



simulator Porta, 1923, Noterus 34

simulatoria Ledoux, Roux & Sawada, 1993, Nebria 93



sinaicus Wewalka, 1984, Hydroporus 61

sinaitica Peyerimhoff, 1907, Coryza 221

sinaitica Schatzmayr, 1936, Tachyura [Tachys] 279

sinaiticus Peyerimhoff, 1907, Chlaenius 354

sinaiticus Alfieri, 1976, Microlestes 424



sinaiticus Schatzmayr, 1936, Scarites 232

sincera Lutshnik, 1933, Amara 547

sincerus Park & Kwon, 1996, Pterostichus 502



sindensis Vazirani, 1969, Laccophilus 76

sindicum Netolitzky, 1935, Bembidion 20, 269

sinensis Peschet, 1915, Acilius 48

sinensis Csiki, 1931, Agonum 454

sinensis Géhin, 1885, Carabus 199

sinensis Chaudoir, 1856, Chlaenius 349



sinensis Wood, 1874, Cicindela 108

sinensis Feng, 1935, Dytiscus 52



sinensis Jedlika, 1957, Harpalus 377

sinensis G. Ochs, 1924, Metagyrinus [Paragyrinus] 28

sinensis Wewalka, 1997, Microdytes 75

sinensis Bates, 1873, Onycholabis 461

sinensis Hope, 1845, Peltodytes [Haliplus] 33

sinensis Jedlika, 1962, Pterostichus 514

sinensis Tschitschérine, 1897, Stenolophus 406

sinensis R. T. Bell & J. R. Bell, 1987, Yamatosa 78

sinester Coiffait, 1959, Aphoenops 294

singalensis H. Kolbe, 1927, Cerapterus 19, 209

singhalensis Andrewes, 1926, Dyschirius 229

singombae J. Schmidt, 1998, Xestagonum 467

singularis Tschitschérine, 1894, Amara 557


singularis Rosenhauer, 1856, Cymindis 414

singularis Tschitschérine, 1906, Harpalus 385

singularis Chaudoir, 1876, Hypaetha [Cicindela] 112

singularis Tschitschérine, 1889, Pterostichus 516

singularis Bates, 1873, Tinoderus [Panagaeus] 448

sinica Hieke, 1997, Amara 551

sinica Yu & Stork, 1991, Amphizoa 35

sinica Fleutiaux, 1889, Myriochila [Cicindela] 114

sinicola Mandl, 1981, Callistomimus 348

sinicola Mandl, 1981, Cicindela 106

sinicola Deuve, 1989, Paragonotrechus [Agonotrechus] 318

sinicus Bates, 1873, Bradycellus 401

sinicus Breuning, 1950, Carabus 189

sinicus Andrewes, 1938, Cillenus 272

sinicus Deuve, 1989, Cychrus 205

sinicus Hope, 1845, Harpalus 385

sinicus Westwood, 1850, Paussus 210

sinicus Casale, 1988, Pseudotaphoxenus 542

sinicus Tschitschérine, 1897, Pterostichus [Feronia] 501

sininensis Semenov, 1898, Carabus 191

sinjanus J. Müller, 1913, Duvalius [Trechus] 306

sinjensis J. Müller, 1906, Laemostenus 534

sinoelongatus Deuve, 1992, Carabus 124

sinomeridionalis Deuve, 1989, Carabus 127

sinomeridionalis Keyimu & Deuve, 1998, Synuchus 546

sinopensis Breuning & Ruspoli, 1970, Carabus 158

sinotibeticola Mandl, 1975, Carabus 147

sintanus Andrewes, 1927, Pterostichus 516



sinuata Motschulsky, 1844, Amara 550

sinuata Panzer, 1792, Cylindera [Cicindela] 110

sinuata Reiche, 1855, Cymindis 413

sinuata Bruneau de Miré, 1964, Nebria 87

sinuata Bates, 1883, Perigona 448

sinuaticollis Puel, 1934, Acinopus 368

sinuaticollis Régimbart, 1899, Agabus 35

sinuaticollis A. Morawitz, 1862, Amara 554

sinuaticollis Habu, 1978, Synuchus 546



sinuaticollis Sloane, 1903, Tachys 275

sinuatipenis Mateu, 1950, Pterostichus [Haptoderus] 492

sinuatocollis Schatzmayr, 1929, Pterostichus 512

sinuatopunctatus Dejean, 1828, Pterostichus 503

sinuatus Tschitschérine, 1893, Harpalus 382



sinuatus Aubé, 1837, Platambus [Agabus] 44

sinuatus Jeannel, 1947, Pterostichus [Haptoderus] 492



sinuatus Motschulsky, 1844, Stenolophus 406

sinuatus Uéno, 1995, Trechiama 323

sinuatus Schaum, 1860, Trechus 341

sinuellus Bates, 1892, Acupalpus 399

sinuosa Ledoux & Roux, 1990, Nebria 93

sioberae Fourès, 1954, Aphoenops 295

sirentensis d'Amore Fracassi, 1908, Calathus 526



sirgueyana Jeannel, 1941, Cicindela 106

sirkungensis Blumenthal & Müting, 1977, Carabus 162

sistani Brouerius van Nidek, 1954, Calomera [Cicindela] 100

sitiens Antoine, 1936, Nebria 18, 85



siziliarum Puel, 1930, Amara 566

sjoelanderi Jedlika, 1965, Bembidion 271

sjoestedti J. Müller, 1935, Dyschirius 230

skarduensis Guignot, 1958, Agabus 39

skiathos Hinterseher, 1981, Ilybius [Agabus] 43

skoazigum Morvan, 1996, Xestagonum 469

skoazkornek Morvan, 1999, Xestagonum 469

skofjeloscensis Daffner, 1996, Anophthalmus 290



skombrosensis Eidam, 1927, Carabus 166

skopini Hieke, 1976, Amara 558



skopljensis Jedlika, 1963, Chlaenius 352

skoraszewskyi Korge, 1971, Bembidion 248

skoupyi Jeanne, 1996, Platyderus 523

skoupyi P. Moravec & Zieris, 1998, Trechus 341

skrimshiranus Stephens, 1828, Stenolophus 406

skutariensis Reitter, 1900, Harpalus 372

skyaphilus Cavazzuti, 1998, Carabus 127

slapniki Drovenik, Mlejnek & J. Moravec, 1997, Orotrechus 317

slavorum Lutshnik, 1915, Pterostichus [Platysma] 507

slivensis Apfelbeck, 1904, Carabus 152

slivkini Kabak, 1994, Poecilus 484

slongoi Daffner, 1989, Orotrechus 317

sloskyi Csiki, 1927, Dyschirius 224

slovacus Hrka & Pulpán, 1980, Duvalius 307

slovenicus Schweiger, 1950, Trechus 334

slovtzovi Mannerheim, 1849, Carabus 149

slovzovi J. R. Sahlberg, 1880, Agabus [Gaurodytes] 36

smaragdiceps Semenov, 1889, Elaphrus 207

smaragdina W. Horn, 1894, Neocollyris [Collyris] 117

smaragdinus Fischer von Waldheim, 1823, Carabus [HN!] 144



smaragdinus G. Hauser, 1921, Carabus [Coptolabrus] 141

smaragdinus Reitter, 1887, Elaphrus 206

smaragdinus Régimbart, 1891, Gyrinus 28

smaragdinus Duftschmid, 1812, Harpalus [Carabus] [HN?] 382

smaragdinus Waltl, 1835, Pogonus 287



smaragdinus Straneo, 1982, Pterostichus 487

smaragdulus Kraatz, 1878, Carabus 171

smaragdulus Fabricius, 1798, Stenolophus [Carabus] 405

smaragdum Andrewes, 1921, Omophron 208



smaragdynum G. Rossi, 1882, Calosoma 120

smetanai Roughley, Xie & Yu, 1998, Amphizoa 35

smetanai Zamotajlov & Sciaky, 1996, Apatrobus 281

smetanai Baehr, 1998, Gunvorita 573

smetanai Farka, 1995, Leistus 81

smetanai Sciaky, 1995, Myas [Trigonognatha] 476

smetanai Uéno, 1995, Queinnectrechus 318

smetanai Deuve, 1988, Stevensius 320

smetanai Kataev & N. Ito, 1999, Trichotichnus 394

smetanai Morvan, 1996, Xestagonum 469

smetanorum R. T. Bell & J. R. Bell, 1989, Yamatosa 78

smirnovi Kryzhanovskij, 1979, Bembidion 244

smithi Kult, 1954, Dyschirius 224

smolikanus Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 305

smyrnense Apfelbeck, 1904, Bembidion 256

smyrnensis Fedorenko, 1996, Dyschirius [Dyschiriodes] 229

smyrnensis Heyden, 1888, Harpalus 382

snizeki Hendrich, 1993, Graptodytes 60

snowdonius Newman, 1833, Agabus [Colymbetes] 39

snowi Bates, 1883, Nebria 94

soazigae Deuve, 1997, Cychrus 206

soazigae Morvan, 1999, Xestagonum 469



sobaekensis Deuve & Remond, 1998, Carabus 142

sobaeksanensis Kwon & Lee, 1984, Carabus 183

sobosanus Habu, 1954, Acupalpus 399

sobrarbensis Lagar Mascaró & Hernando, 1987, Hydraphaenops 311

sobrina Gory, 1833, Cylindera [Cicindela] 109



sobrina Péringuey, 1896, Drypta 358

sobrina Schaufuss, 1862, Nebria 87

sobrinum Boheman, 1848, Bembidion 254

sobrinus Ménétriés, 1832, Carabus 151

sobrinus Jeannel, 1926, Duvalius 307



sobrinus Dejean, 1829, Harpalus 381

sobrinus Dejean, 1826, Pheropsophus [Brachinus] 218

sochondensis Shilenkov, 1999, Nebria 89

socia Schaum, 1863, Tachyura [Tachys] 277

socialis Schaum, 1864, Zabrus 571

sociennus J. Balfour‑Browne, 1952, Copelatus 48

socius Knirsch, 1929, Duvalius [Duvaliotes] 300

socius Apfelbeck, 1902, Trechus 326

sodalicius Heyden, 1885, Pterostichus 501



sodalis Duftschmid, 1812, Badister [Carabus] 441

sodalis Lapouge, 1925, Carabus 146

sodalis Csiki, 1932, Harpalus 380

sodalis Jeannel, 1960, Trechus 341



sodata Escalera, 1914, Cicindela 106

soederbomi Jedlika, 1965, Bembidion 257



soederbomi Liebke, 1935, Cymindis 415

soederbomi Jedlika, 1935, Poecilus [Pterostichus] 484



soganlica Jedlika, 1965, Nebria 86

soganlidaghensis Lassalle & P. Machard, 1991, Carabus 163

soganliensis Heinz & Korge, 1964, Carabus 181

sogdiana Kryzhanovskij, 1974, Amara [Curtonotus] 561

sogdiana Semenov, 1904, Cicindela 104

sogdianum Belousov & Mikhailov, 1990, Bembidion 262

sogdianus Semenov, 1898, Carabus 153

sogdianus Putchkov & Dolin, 1998, Leistus 84

sogdianus A. I. Jakovlev, 1897, Platambus [Anagabus] 44

sogdianus Belousov & Kabak, 1998, Trechus 341



soginoides J. R. Sahlberg, 1903, Harpalus 372

sohei Pawowski, 1974, Tachys 275

sojaki Kirschenhofer, 1987, Pterostichus 505

sokhensis Kabak, 1998, Poecilus 484



sokolari Born, 1904, Carabus 152

sokolari Roubal, 1910, Omophron 208

sokolovi Zamotajlov, 1988, Deltomerus 283

sokolovi Ljovuschkin, 1963, Duvalius 305

sokolovi Belousov, 1990, Trechus 341

sokolowskii Fassati, 1957, Bembidion 271

solanum Jedlika, 1965, Bembidion 265



solarii Baliani, 1943, Amara 564

solarii J. Müller, 1918, Bembidion 268



solarii Gestro, 1898, Duvalius [Anophthalmus] 302

solarii Schatzmayr, 1929, Pterostichus 503

solarii Magrini & Sciaky, 1995, Rhegmatobius 240

solarii Jeannel, 1921, Trechus 341

solaris Gistel, 1857, Harpalus 387



solata Coquerel, 1859, Amara 547

solers Sturm, 1824, Pterostichus [Platysma] 508

soleymanensis Morvan, 1974, Ocys 273



soleymanensis Morvan, 1976, Trechus 329

solidior Deuve & Imura, 1990, Carabus 127

solidior Uéno, 1975, Ryugadous 319

solidus Lapouge, 1899, Carabus 168

solieri Aubé, 1837, Agabus 39

solieri Putzeys, 1870, Amara 565


solieri Bassi, 1834, Calathus 528

solieri Dejean, 1826, Carabus 140



solieri Dejean, 1831, Harpalus 370

solieri Aubé, 1838, Hyphoporus [Hydroporus] 73



solinghoensis Jedlika, 1935, Chlaenius 348

solissimus Antoine, 1939, Platyderus [Iblania] 523



solitaria Gistel, 1857, Amara [Celia] 557

solitaris Dejean, 1829, Harpalus [NP] 382

solitarius Ledoux & Roux, 1999, Archastes 79

solitarius Wollaston, 1863, Nesacinopus [Cratognathus] 388

solitarius Peyron, 1858, Penthophonus [Ditomus] [DA] 393

solivagus Péringuey, 1926, Chlaenius 351

sollaudi Jeannel, 1916, Trichaphaenops [Trechus] 344

sollicita Pantel, 1888, Amara 558

sollicitans Hartert, 1907, Carabus 166

solodovnikovi Zamotajlov, 1992, Carabus 132

solskii A. I. Jakovlev, 1897, Agabus 40

solskyanum Obydov, 1999, Calosoma [Callisthenes] 119

solskyi Heyden, 1880, Amara [Leiocnemis] 558

solskyi Reiche, 1864, Amblystomus 360

solskyi Netolitzky, 1934, Bembidion 262



solskyi Zaitzev, 1918, Calosoma 119

solskyi Ballion, 1878, Carabus 145

solskyi Tschitschérine, 1896, Cymindis 416

solskyi Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403

solskyi Komarov, 1995, Metablus 431



solskyi Zaitzev, 1916, Omophron 208

solskyi Chaudoir, 1878, Pterostichus [Feronia] 513



solstitialis Gistel, 1837, Calomera [Cicindela] 100

solus Leech, 1949, Agabus 38

solus Cavazzuti, 1997, Cychrus 205

soluta Dejean, 1822, Cicindela 107

solutus Oberthür, 1885, Carabus 168

solutus Sharp, 1882, Laccophilus 76

soma Mateu & Deuve, 1979, Trechus 341



somalicum Alluaud, 1935, Omophron 208

somcheticus Schauberger, 1933, Harpalus 370

somervelli Andrewes, 1930, Amara 550

sommeri Mannerheim, 1844, Carabus 185



somnolenta Putzeys, 1866, Amara [Curtonotus] 561

somoni Jedlika, 1968, Amara 561



somoni Jedlika, 1967, Cymindis 415

sonamargensis Deuve, 1981, Carabus 155

sonamargensis Balkenohl, 1994, Dyschirius 229

sonei Uéno, 1992, Kurasawatrechus 313



sonei Uéno, 1982, Trechiama 323

songarica Putzeys, 1866, Amara [Amathitis] 562



songaricus Motschulsky, 1850, Carabus 136

songinus Jedlika, 1968, Poecilus [Pterostichus] 481

songorica Motschulsky, 1845, Cicindela 107

songoricus Gebler, 1859, Agabus [Colymbetes] 38

songoricus Kirsch, 1859, Carabus 145

songoricus Motschulsky, 1845, Pterostichus [Molops] 506

songoricus Belousov & Kabak, 1992, Trechus 341

songpanensis Deuve, 1998, Amerizus 242

songpanensis Deuve, 1991, Cychrus 205

songpanicus Deuve, 1990, Carabus 191

songshanicus Kleinfeld, 1994, Carabus 127

sonkulensis Vereschagina & Kabak, 1997, Laemostenus 537

sontiacus J. Müller, 1935, Anophthalmus 292



sonticus Bernau, 1914, Carabus 184

sonuta Uéno, 1992, Trechiama 323

soosi Csiki, 1912, Anophthalmus 292

sorakensis Deuve, 1998, Carabus 142

sordidatus Habu, 1954, Trephionus 547

sordidum Dejean, 1828, Agonum 451

sordidus Andrewes, 1933, Brachinus 216



sordidus Herbst, 1784, Hydroporus [Dytiscus] 64

sordidus Sharp, 1882, Hydrovatus 70

sordidus Sharp, 1882, Laccophilus 77



sordidus Marsham, 1802, Paranchus [Carabus] 462

soreli Schuler, 1964, Amara [Celia] 557

sorianus Jeanne, 1972, Carabus 167

sorini Deuve & Mourzine, 1997, Cychrus 204

soriscensis Daffner, 1996, Anophthalmus 292

soror W. Horn, 1897, Cylindera [Cicindela] 111



soror K. Daniel, 1903, Oreonebria [Nebria] 95

soror Rambur, 1837, Singilis 438



sororcula Tschitschérine, 1895, Amara 547

sororius A. Morawitz, 1886, Carabus 146



sorredensis F. Reymond & A. Reymond, 1935, Carabus 130

sotai Ishikawa & Kubota, 1994, Carabus 178

sotilloi Español, 1971, Hypotyphlus 239

sotkaensis Jedlika, 1958, Pterostichus 508

sotshiensis Zamotajlov, 1997, Lindrothius 529

sotshiensis Belousov, 1987, Trechus 341



souletinus Jeanne, 1970, Zabrus 569

sousae Vuillefroy, 1868, Pterostichus [Feronia] 497

soussieuxi Perreau & Quéinnec, 1987, Geotrechus 310

spadiceus Dejean, 1828, Pterostichus [Feronia] 492


spaethi Ganglbauer, 1904, Duvalius [Trechus] 305

spaethi Munster, 1922, Notiophilus 96

spaethi Reitter, 1913, Notiophilus 97

spagnoloi Gestro, 1885, Duvalius [Anophthalmus] 303

sparreschneideri Munster, 1922, Haliplus 32

sparsepilifer Uéno, 1975, Oroblemus 316



sparsus Marsham, 1802, Noterus [Dytiscus] 34

sparsutus Reitter, 1898, Eucarterus [Eriotomus] 366

spartanum Fassati, 1944, Bembidion 263

spartanus Schaum, 1862, Molops [Pterostichus] 475

spasskianus Fischer von Waldheim, 1823, Carabus 174

spathulifer Bates, 1873, Chlaenius 353

spatiosus Andrewes, 1932, Apristus 431

speciense Jedlika, 1932, Bembidion 271



spectabilipennis Deuve, 1989, Carabus 123

spectabilis Schaum, 1858, Amara 563

spectabilis Joseph, 1871, Anophthalmus 293

spectabilis Csiki, 1906, Carabus 173

spectabilis Hampe, 1852, Zabrus 572

spectans Jedlika, 1933, Bembidion 271

spectus Jedlika, 1962, Poecilus [Pterostichus] 484



speculare Küster, 1847, Bembidion 253

specularis Chaudoir, 1865, Myriochila [Cicindela] 114

speculator Harold, 1878, Agonum [Colpodes] 454

speculifer Andrewes, 1929, Dyschirius 229

speculifera Chevrolat, 1845, Myriochila [Cicindela] 114

speiseri Ganglbauer, 1892, Duvalius [Trechus] 300

speiseri Ganglbauer, 1891, Nebria 87

spelaeophila Obenberger, 1917, Nebria 85

spelaeus Reitter, 1870, Duvalius [Trechus] 307

spelaeus Uéno, 1958, Kurasawatrechus 313

speleus Cobos, 1961, Platyderus 523

speluncarius Reitter, 1916, Neotrechus [Anophthalmus] 315

speluncarum Csiki, 1912, Duvalius [Anophthalmus] 299

speluncicola Chaudoir, 1868, Speluncarius [Feronia] 516

spencei Gistel, 1838, Glycia [Cymindis] 409

spencii Stephens, 1828, Bembidion [Lopha] 248

spengeli G. Hauser, 1921, Carabus [Coptolabrus] 143

sphaericus DeGeer, 1774, Hyphydrus [Dytiscus] 75

sphaerithorax Ledoux & Roux, 1999, Nebria 89

sphaerocephalus Olivier, 1795, Dixus [Scarites] 365

sphaerulifer Bates, 1873, Dyschirius 229

sphex Andrewes, 1933, Calodromius [Dromius] 419

sphinx Reitter, 1895, Carabus 149

sphodriformis Bates, 1873, Pterostichus 498


sphodrinus Fischer von Waldheim, 1844, Carabus 181

sphodrinus Gautier des Cottes, 1866, Carabus 186

sphodroides Wollaston, 1862, Calathidius [Calathus] 532

spiculifer Bates, 1883, Pterostichus 494

spiessi Jeannel & Mallász, 1929, Duvalius 300

spilotus Illiger, 1798, Calodromius [Carabus] 419

spilotus Dejean, 1825, Syntomus [Dromius] 433

spilotus Bates, 1886, Tachys 276

spinangulus Reitter, 1913, Leistus 18, 84

spinibarbis Fabricius, 1775, Leistus [Carabus] 84



spinicollis Breit, 1913, Aphaenopsis [Anophthalmus] 293

spinicollis L. Dufour, 1857, Cychrus 205



spinicollis Chaudoir, 1843, Ditomus [Ditomus] 365

spinicollis A. Gaudin, 1947, Geotrechus 310

spinicollis Dejean, 1828, Molopidius [Feronia] 472

spinifer Jeannel, 1928, Duvalius 300

spinifer Schaufuss, 1862, Laemostenus [Pristonychus] 539

spinifer Kasahara, 1994, Trichotichnus 397

spinifera Antoine, 1939, Cymindis 413

spiniger Paykull, 1790, Sphodrus [Carabus] 543

spinigera Eschscholtz, 1829, Cicindela 106



spinilabris Panzer, 1797, Leistus [Carabus] 82

spinipes Fischer von Waldheim, 1844, Acinopus 369

spinipes Fabricius, 1798, Zabrus [Blaps] 572

spiniplatus Belousov & Kabak, 1998, Trechus 335

spinolae Cristoforis & Jan, 1837, Carabus 158

spinolae Gestro, 1889, Cylindera [Cicindela] 112

spinolae Dejean, 1828, Pterostichus [Feronia] 504

spinosa Ledoux & Roux, 1995, Nebria 95


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