Associations of shrimp with gobies are widespread across the tropics. Most of the work on these goby-shrimp relationships has been done in the Red Sea/Indian Ocean Luther 1958; Magnus 1967; Karplus 1981

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Figure 6

Figure 7

Sample ID #	Date of collection	Months after exclosure was constructed
0	February 23-27	0
------------	February 28	Exclosure Constructed
2	April 26-30	2
3	May 21-25	3
4	June 21-26	4
5	August 11-13	5.5
------------	August 15th	Exclosure taken down
6	September 13-15	6.5
7	October 5-6	7

Table 1. Dates of data collection, corresponding sample ID numbers, and timing relative to exclosure events.

Time	Burrow maintenance	Foraging	Static Outside	Total time outside
600	13.1 + 10.2	27.0 + 8.4	15.6 + 5.6	55.1 + 13.5
900	4.8 + 13.4	17.2 + 3.5	17.2 + 4.6	38.9 + 16.3
1200	44.7 + 12.8	20.4 + 10.1	18.9 + 7.2	83.1 + 21.3
1500	74.9 + 11.1	45.7 + 12.2	58.4 + 15.9	178.6 + 21.3
1800	108.9 + 16.6	56.4 + 10.3	26.0 + 21.9	191.3 + 26.2

Table 2. Duration of behaviors in seconds (mean + SD) recorded for shrimp during five 1000-sec sampling periods in daylight.

Status of the goby	‘Bout’ length (Time a shrimp spent out of the burrow – in seconds)	Percent of time shrimp were out of the burrow	Percent of shrimp emergences when ‘burrow maintenance’ occurred
Present (N=7)	14.0+6.4	53.6+21.8	52.7+25.1
Absent (N=7)	3.1+0.2	6.9+3.4	98.7+2.8

Table 3. Means (+ S.D.) from seven shrimp burrows with associated goby and seven burrows without: ‘bout’ length; percent of total time that shrimp were out of the burrow; percent of shrimp emergences that involved burrow maintenance.
.

Source	df	SS	MS	F	P
Treatment	2	14.99	7.49	1.07	0.345
Month	5	338.74	84.69	12.08	< 0.001
Interaction	8	28.01	3.50	0.500	0.856
Error	255	1788.06	7.01
Total	269	2169.8

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