Atlantis operating model equations



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2.4 Growth of consumers

The growth of an invertebrate consumer (CX) is given by:



(C.47)

with CX the growth efficiency of CX when feeding on live prey, CX,j the efficiency when feeding on detritus (DL treated separately to DR), space limitation given by:



(C.48)

where CXmax is the maximum biomass per area allowed for CX, CXlow is the crowding lower threshold, CXsat is the crowding half saturation level, CXthresh is the crowding threshold (this formulation is based on that of the European Regional Seas Ecosystem Model II (ERSEM II) (Blackford 1997)), substrate is the proportion of the available space that is of the correct substrate type to support this type of primary producer and habdegrad is the scalar for local habitat degradation scenarios (see equation 4.1 in the main text for the equation used).



The oxygen limitation is given by:

(C.49)

where o2 is the depth of the oxygen horizon and CX,Mo2 is the half oxygen mortality depth.



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