Leafcutter and Mason Bees: a Biological Catalogue of the Genus Megachile of the Neotropics Anthony Raw



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Subgenus Schrottkyapis Mitchell


Eumegachile subgenus Schrottkyapis Mitchell 1980: 46. Type species: Megachile assumptionis Schrottky 1908: 233 (not 1913a: 153). Original designation and monotypy.

The subgenus comprises a single species.


Megachile (Schrottkyapis) assumptionis Schrottky

Megachile assumptionis Schrottky 1908 (April): 233. Female and male. Type locality: Asuncion, Paraguay, 22 January 1906 (Anisits). Type repository: MZSP ?

Megachile armigera Friese 1908a (May): 69. Female. Type locality: Villa Rica, Paraguay (Burgdorf). Type repository: ? Synonymy of Schrottky 1913a: 153.

Megachile (Sayapis) assumptionis Mitchell 1943b: 664.

Eumegachile (Schrottkyapis) assumptionis Mitchell 1980: 46.

ADDITIONAL REFERENCES. Almeida et al 1997: 344-352 (F, N). Friese 1908a 63 (K). Martins & Dealmeida 1994 (N). Schrottky 1913a: 136 & 146 (K). Silveira et al 2002: 214 (D). Raw coll.

DISTRIBUTION. BRAZIL: DF; MG. PARAGUAY: Asuncion, Villarrica.


Subgenus Stelodides Moure


Megachile subgenus Stelodides Moure 1953: 123-124. Type species: Megachile euzona Pèrez. Original designation and monotypy.

Chalicodoma subgenus Stelodides Michener 1962: 20.

Chrysosarus subgenus Stelodides Mitchell 1980: 72.

The subgenus comprises a single species.


Megachile (Stelodides) euzona Pèrez

Megachile euzona Pèrez 1899: 105-106. Male. Type locality: Valparaiso, Chile (M. Porter). Type repository: MNHN.

Megachile philippii Friese 1905: 139-140. Male & female ? Type locality: Santiago, Chile (Philippi). Type repository: ? Synonymy of Vachal 1909: 8.

Anthidium euzona Friese 1925: 42.

Anthidium philippi Friese 1925: 42.

Megachile (Stelodides) euzona Moure 1953: 124 (D).

Chrysosarus (Stelodides) euzona Mitchell 1980: 72.

ADDITIONAL REFERENCES. Claude-Joseph 1926: 254-255 (L, M, N).

DISTRIBUTION. CHILE: Aconcagua, Hacienda Las Mercedes; Pemehue (1650m); San Bernardo; Santiago.

NESTING. Nest in old beetle galleries in wood. Cuts petals of Althaea rosea, Clarkia pulchella and Lavatera grandiflora.

FLOWERS. Alstroemeria, Cichorium intybus, Hibiscus and Medicago sativa (alfalfa).

LIFE HISTORY. Development lasts six weeks. Prepupa overwinters in cocoon. Adults fly in November and December.

MORPHOLOGY. Larva described.


Subgenus Trichurochile Mitchell


Cressoniella subgenus Trichurochile Mitchell 1980: 63. Type species: Megachile thygaterella Schrottky. Original designation and monotypy.

Megachile (Trichurochile) Michener 2000: 566.

The three members of this subgenus range from northern Argentina to central Brazil.




  1. Megachile (Trichurochile) gracilis Schrottky

Megachile gracilis Schrottky 1902: 435-436. Female. Type locality: Jundiay [=Jundiaí], São Paulo, 28 January 1900 (Beron). Type repository: MZSP. Examined. [Type missing fore tibiae and tarsi, right mid tibia and tarsi and left hind tarsi.]

Megachile (Trichurochile) gracilis Raw 2002 #7: 35.

ADDITIONAL REFERENCES. Ihering 1904: 470 (N). Schrottky 1913a: 138 (K). Silveira et al 2002: 214 (D).

DISTRIBUTION. BRAZIL: MG; SP.

NESTING. Nest lined with cut leaves. The cell is 12 mm long and 5 mm wide.




  1. Megachile (Trichurochile) lachnopyga Moure

Megachile lachnopyga Moure 1941: 94-96. Male. Type locality: Curitiba (900 m), Paraná [Brazil]. Type repository: Claretiano collection. Synonymy of Moure 1943: 186.

Megachile lachnopyga var. langei Moure 1941: 96. Male. Type locality: Rio Baraguí, Curitiba, Paraná (Rodolfo Lange). Type repository: Claretiano. Synonymy of Moure 1943: 186.

ADDITIONAL REFERENCE. Silveira et al 2002: 214 (D).

DISTRIBUTION. BRAZIL: PR- Curitiba; SP.


  1. Megachile (Trichurochile) thygaterella Schrottky

Megachile thygaterella Schrottky 1913a: 220-221. Male. Type locality: Puerto Bertoni, Paraguay. Type repository: MZSP.

Megachile anthidioides Schrottky 1919 (in part). Synonymy of Moure 1943: 186.

Cressoniella (Trichurochile) thygaterella Mitchell 1980: 63.

ADDITIONAL REFERENCE. Silveira et al 2002: 214 (D). RAW.

DISTRIBUTION. BRAZIL: DF- Brasilia; SP. PARAGUAY: Puerto Bertoni.


Subgenus Tylomegachile Mitchell


Megachile subgenus Tylomegachile Moure 1953: 120-121. Type species: Megachile orba Schrottky. Original designation and monotypy. Michener et al 1994: 149, Michener 2000: 566.

Cressoniella subgenus Tylomegachile Mitchell 1980: 63.

The six species at present included in the subgenus occur in widespread localities: Mexico, Costa Rica, St. Vincent (West Indies), Colombia, Peru and Paraguay.




  1. Megachile (Tylomegachile) eulaliae Cockerell

Megachile eulaliae Cockerell 1917b: 239-240/ Cockerell 1919a: 218-219. Female. Type locality: Santa Eulalia, Peru, 18 January 1913 (C. H. T. Townsend). Type repository: USNM 21689. Examined.

Megachile (Tylomegachile) eulaliae Raw 2002 #7: 35.

DISTRIBUTION. PERU.




  1. Megachile (Tylomegachile) hamata Mitchell

Megachile hamata Mitchell 1930: 295-296. Female. Type locality: Rio Frio, Magdalena, Colombia, 26 December 1926 (Salt). Type repository: NHML 17a2472. Examined.

Megachile (Tylomegachile) hamata Raw 2002 #7: 35.

DISTRIBUTION. COLOMBIA: Magdalena- Rio Frio, Santa Marta.

FLOWERS. Machaerium humboldtiarum.


  1. Megachile (Tylomegachile) orba Schrottky

Megachile orba Schrottky 1913a: 214. Male. Type locality: Puerto Bertoni, Paraguay. Type repository: MZSP.

Megachile pseudocoelioxys Schrottky 1920: 28-29. Female. Type locality: Franca, São Paulo state, Brazil, January 1903. Type repository: MZSP. Examined. Synonymy of Moure 1953: 121.

Megachile nuda Mitchell 1930: 296-296. Female. Type locality: Pedra Branca, Brazil. Type repository: ANSP 4130. Synonymy of Moure 1953: 121.

(Not M. nuda Rayment 1935. Australia. [= Chalicodoma (Hackeriapis) nuda]. Designation of Michener 1965: 198.)



Megachile (Tylomegachile) orba Moure 1953: 121.

Megachile (Ptilosarus) orba Michener 1954: 102 (D, F, L). (Erroneous listing).

Cressoniella (Tylomegachile) orba Mitchell 1980: 63.

ADDITIONAL REFERENCES. Mitchell 1930: 255 (D). Silveira et al 2002: 214 (D). RAW.

DISTRIBUTION. ARGENTINA: MI- Iguazú. BOLIVIA: Buenavista. BRAZIL: BA- Salvador; DF- Brasilia; MG; MS- Corumbá, Pedra Branca; MT- Bodoquena, Camisão, Chapada dos Guimarães; PR- Curitiba, Iguaçú; RN; RS; SP- Batatais, Franca, Guarulhos, Onda Verde, Rio Claro, São Paulo. COLOMBIA: Magdalena- Rio Frio. PANAMA: Canal Zone- Juan Mina. PERU: Tingo Maria (670 m).

FLOWERS. Cornuta grandiflora, Machaerium humboldtiarum.

LIFE HISTORY. In Panamá adults fly in June.


  1. Megachile (Tylomegachile) simplicipes Friese

Megachile simplicipes Friese 1921: 93. Male & Female ? Type locality: San José, Costa Rica, 1913 (Schmidt). Type repository: ?

Megachile nigrolateralis Cockerell 1919: 217-218. Female. Type locality: Mexico (Baker). Type repository: USNM 21688. Synonymy of Mitchell 1930: 295.

(Not M. lachesis nigrolateralis Cockerell 1914a: 279. Phillippines [= Creightonella lachesis (Smith) 1861].



Megachile atrolateralis Cockerell 1927c: 5. New name for M. nigrolateralis Cockerell 1914a).

Cressoniella (Tylomegachile) simplicipes Mitchell 1980: 63.

Megachile (Tylomegachile) simplicipes Ayala et al 1997: 455 (D).

ADDITIONAL REFERENCE. Cockerell 1949: 450 (D).

DISTRIBUTION. MEXICO: CH; JA; MO; NA; OA; SI. COSTA RICA. HONDURAS: Zomorano.

LIFE HISTORY. April to July.




  1. Megachile (Tylomegachile) toluca Cresson

Megachile toluca Cresson 1878: 129. Male. Type locality: Mexico (Sumichrast). Type repository: ANSP 2432.

Megachile (Tylomegachile) toluca Ayala et al 1997: 455 (D).

ADDITIONAL REFERENCE. Cresson 1916: 132 (T).

DISTRIBUTION. MEXICO: JA; ME; NA; OA; PU; SO; VC.


  1. Megachile (Tylomegachile) tridentata Ashmead

Megachile tridentata Ashmead 1900: 213. Male. Type locality: St. Vincent (?) [West Indies]. Type repository: NHML 17a2424. Examined.

Megachile (Tylomegachile) tridentata Raw 2002 #7: 35.

Ashmead (1900) states he is "not certain whether it was taken on this island or on Grenada."

ADDITIONAL REFERENCE. Friese 1908a: 38 (D).

DISTRIBUTION. ST. VINCENT.





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