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References


Altringham, J.D., O'Brien, S. & Julian, S. (1994) Feeding ecology of the little owl (Athene

noctua) an an upland site in northern England. Naturalist 119: 81-94.

Bevan, G. (1964) The food of tawny owls in London. London Bird Report 29: 56-72. Brown, R., Ferguson, J., Lawrence, M. & Lees, D. (1987) Tracks and Signs of the Birds of Britain and Europe. Christopher Helm, London.

Macdonald,A.W.(1976) Nocturnal observations of tawny owls (Strix aluco) preying upon

earthworms. Ibis 118: 579-80.

Mead, C. (1987) Owls. Whittet, London.

Southern, H.N. (1954) Tawny owls and their prey. Ibis 96: 384-410.

Wardhaugh, A.A. (1983) Owls of Britain and Europe. Blandford, Poole.

Yalden, D.W. (1977) The Identification of Remains in Owl Pellets. The Mammal Society,London,



Table 1:Monthly Record of Pellets Collected and Presence of Owls at Roost



Collection Date

No. of Pellets

Volume (cm3)

No. of Owls Seen












24. 1. 93

17

-

1

19. 2. 93

12

-

2

27. 3. 93

16

127

0

21. 4. 93

20

138

0

22. 5. 93

14

77

0

21. 6. 93

7

39

0

22. 7. 93

16

104

0

22. 8.93

0

0

0

25. 9. 93

1

5

0

25.10.93

1

4

2

20.11. 93

6

38

2

20.12. 93

14

82

2

22. 1. 94

7

29

1

20. 2. 94

6

30

0

26. 3. 94

32

187

1

24. 4.94

15

103

0

23. 5. 94

1

6

0

22. 6.94

1

3

0

25. 7.94

1

6

0

22. 8. 94

0

0

0

25. 9.94

0

0

0

26.10.94

0

0

0

19.11.94

0

0

0

22.12. 94

1

6

0

TOTAL

159

984

-


Table 2:Monthly Record of Prey Items

Collection Date

Bank Vole

Short-tailed Vole

Wood Mouse

Common Shrew

Pygmy Shrew

Other

24. 1. 93

26

2

18

0

1

1 bird

19. 2.93

12

0

5

0

0




27. 3. 93

14

4

8

1

0




21. 4. 93

9

5

4

2

1

1 probable rat

22. 5.93

4

5

2

3

0




21. 6.93

2

3

2

0

0




22. 7.93

9

2

3

2

0




22. 8.93

0

0

0

0

0




25. 9. 93

1

0

0

0

0




25.10.93

1

0

0

0

0

1bird

20.11. 93

7

2

0

0

0




20.12. 93

7

1

8

2

0




22. 1.94

4

0

1

0

0




20. 2. 94

3

2

1

0

0




26. 3. 94

4

11

6

2

4

1 bird (robin?)



















1 young rabbit

24. 4. 94

4

6

1

2

0

1 probable



















young rabbit*

23. 5.94

0

0

0

0

0




22. 6. 94

0

1

0

0

0




25. 7.94

1

0

0

0

0




22. 8.94

0

0

0

0

0




25. 9.94

0

0

0

0

0




26.10. 94

0

0

0

0

0




19.11.94

0

0

0

0

0




22.12.94

2

0

0

0

0




TOTAL

111

43

59

14

6

rat 1, rabbit 2



















mole 2, bird 3

NO. PER100 cm3 **

7.42

4.17

3.66

1.42

0.61



*Remains of rabbits found in pellets collected on 26.3.94 and 24.4.94 could be from the

same individual but here counted as two.

** For March 1993 to December 1994 inclusive. (Pellet volumes not recorded January and February 1993).


Total number of prey items identified: 241

Number of prey items per 100cm3 (all species): 17.99


Table 3: Summary of Prey Items Identified

Species

Number of Individuals




Mass Represented*







(%)

g

(%)

Bank vole

111

(46.1)

1776

(43.6)

Short-tailed vole

43

(17.8)

903

(22.1)

Wood mouse

59

(24.5)

1062

(26.0)

Common shrew

14

(5.8)

112

(2.7)

Pygmy shrew

6

(2.5)

24

(0.6)

Mole

4

(0.8)

140

(3.4)

Probable Rat

1

(0.4)

60

(1.4)

Probable Rabbit

2

(0.8)

-

-

Bird

3

(1.2)

-

-
















Total

241

4077

-

-

* Based on figures for mean mass of each prey species provided in Yalden (1977)


For rabbit and birds there were insufficient remains to estimate size and therefore mass

hence these species were not included in the calculation of percentage mass.

Table 4:Prey Items Identified in Tawny Owl Pellets from elsewhere in the Cleveland Area

Site

Date

Prey Items

Comments












Cross Wood, Flatts Lane

(NZ 543169)



Spring

1967


short-tailed vole 1

bank vole 3

bird 1


Roost site approx. 0.6 km NW of site of present study; beech tree cavity. Unrecorded small no. of pellets.












Wood at Ormesby Hall

(NZ 530171)



5.4.1971

wood mouse 4

common shrew 1

house sparrow 10


Roost in a yew tree. Unrecorded small no.of pellets.













Plantation near incline, Ingleby Greenhow

(NZ 602025)



2.4.1978

short-tailed vole 2

bird 5


Roost in a holly tree. Unrecorded small no. of pellets.













Wood at Teesside High School

(NZ 425149)



9. 6.93

birds 2

48 cm3 of pellets




5. 7.93

birds 4

50cm3 of pellets




6.12.93

wood mouse 4

50 cm3 of pellets







brown rat 2

Pellets collected from a variety of sites within the wood



.

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