Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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formosanus N. Ito, 1992, Anisodactylus 361

formosanus Jedlika, 1939, Brachinus 215



formosanus Uchida & Tamanuki, 1927, Carabus [Acoptolabrus] 148

formosanus Habu, 1965, Chlaenius 353

formosanus Jedlika, 1935, Chlaenius 356

formosanus Tanaka, 1958, Chydaeus 362

formosanus Dupuis, 1912, Cillenus 272



formosanus N. Ito, 1993, Coleolissus 369

formosanus Jedlika, 1940, Colpoides 457

formosanus Jedlika, 1939, Craspedophorus 446

formosanus Jedlika, 1940, Dromius [Lebidromius] 420

formosanus Kult, 1949, Dyschirius 225

formosanus Jedlika, 1940, Holcoderus 436

formosanus Jedlika, 1954, Kaszabellus 459

formosanus Kamiya, 1938, Lacconectus [Platynectes] 48



formosanus Takizawa, 1932, Laccophilus 76

formosanus Jedlika, 1952, Microlestes 422

formosanus Jedlika, 1940, Myas [Trigonognatha] 476

formosanus Takizawa, 1931, Orectochilus 28

formosanus Jedlika, 1940, Parophonus [Hyparpalus] 392

formosanus Dupuis, 1913, Pericalus 437

formosanus Dupuis, 1912, Pheropsophus 217

formosanus Dupuis, 1914, Pheropsophus [HN] 217

formosanus Jedlika, 1939, Planetes 359



formosanus Dupuis, 1912, Platymetopus [Dioryche] 393

formosanus Jedlika, 1939, Platynus [Colpodes] 464

formosanus Jedlika, 1956, Pogonus 287

formosanus Kamiya, 1938, Rhantus 46

formosanus Lindroth, 1956, Synuchus 545

formosanus Jedlika, 1949, Trichotichnus 395

formosensis Lorenz, 1998, Chlaenius 353

formosula Antoine, 1939, Cymindis 414

formosum Andrewes, 1935, Asaphidion 242



formosum C. R. Sahlberg, 1827, Bembidion 244

formosus Semenov, 1887, Carabus 142



formosus Chaudoir, 1856, Chlaenius 354

formosus Wollaston, 1867, Copelatus 47

formosus Habu, 1954, Morimotoides 460

formosus Wasmann, 1912, Paussus 210

formosus Semenov, 1895, Reflexisphodrus [Taphoxenus] 543

formosus Aubé, 1838, Stictonectes [Hydroporus] 69

formosus Semenov, 1889, Stomis 517

formosus Schauberger, 1935, Trichotichnus 395

fornicatum Beck, 1817, Bembidion 247

fornicatus Kolenati, 1845, Pterostichus [Omaseus] 495

forojulensis Busulini, 1959, Orotrechus 316

forsteri Mandl, 1981, Carabus 138

fortepunctata Eichler, 1924, Tachyura [Tachys] 279

fortepunctatus S. Battoni & Blumenthal, 1973, Carabus 157

fortepunctatus Schauberger, 1934, Tschitscherinellus 367

fortestriatum Mannerheim, 1852, Bembidion 248

fortestriatus Poppius, 1906, Pterostichus [Cryobius] 492


fortetuberculatus Heinz, 1983, Carabus 155

forticornis Fairmaire, 1888, Steropanus 517



forticostis Kraatz, 1878, Carabus 136

forticostis Kraatz, 1897, Carabus [Chrysocarabus] 140

fortii J. Müller, 1963, Orotrechus 317

fortimanus Reitter, 1903, Trechus 331

fortipes Chaudoir, 1850, Poecilus [Feronia] 482

fortipes Uéno, 1999, Trechus 331

fortis A. Morawitz, 1862, Pterostichus 509

fortiterpunctatus Guignot, 1932, Graptodytes 59

fortius Netolitzky, 1920, Bembidion 269

fortunatum Wollaston, 1871, Bembidion 253

fortunatus Heinz, 1970, Carabus 163

fortunatus Wollaston, 1863, Nesacinopus [Cratognathus] 388

fortunatus Mateu, 1957, Philorhizus 425

fortunatus Machado, 1989, Tachys 275

fortunatus Jeannel, 1927, Trechus 331

fortunei A. Adams, 1861, Carabus 148



fossarum Germar, 1824, Agabus [Dytiscus] 36

fossifrons Putzeys, 1866, Dyschirius 228

fossiger Chaudoir, 1877, Carabus 200

fossiger Kraatz, 1897, Carabus [Platycarabus] 184

fossiger Jedlika, 1937, Pterostichus 506

fossor Linné, 1758, Clivina [Tenebrio] 219



fossor Fabricius, 1781, Zabrus [Tenebrio] 573

fossulatus Dejean, 1826, Carabus 199

fossulatus Quensel, 1806, Pterostichus [Carabus] 487

fossulicola Antoine, 1943, Dyschirius 229

fossulifer A. Fleischer, 1893, Carabus 151

fossulipennis W. J. MacLeay, 1871, Hyphydrus [Hydroporus] 74

fosteri Aguilera & Ribera, 1996, Deronectes 58

foucauldi Bruneau de Miré, 1958, Laemostenus [Sphodroides] 539

foudrasi Dejean, 1829, Carabus 187

foudrasii Dejean, 1826, Nebria 86

fouresi A. Gaudin, 1945, Hydraphaenops 311

foveata Gebler, 1829, Blethisa 206

foveatipennis Csiki, 1927, Carabus 138

foveatus G. Hauser, 1913, Carabus [Apotomopterus] 126

foveatus Geoffroy, 1785, Syntomus [Buprestis] 432



foveibasis Jedlika, 1956, Amara 558

foveicollis Schaufuss, 1882, Carabus 163

foveicollis Putzeys, 1861, Clivina 220

foveicollis Motschulsky, 1844, Harpalus 381

foveicollis N. Ito, 1998, Trichotichnus 394

foveiger Tschitschérine, 1895, Harpalus 376


foveipennis Lebis, 1926, Carabus [Chrysocarabus] 139

foveipennis Casale, 1988, Platyderus [Pseudotaphoxenus] 522

foveipennis Heyden, 1883, Zabrus 570

foveocollis Eschscholtz, 1823, Patrobus [Platysma] 285



foveola Gyllenhal, 1810, Syntomus [Lebia] 432

foveolata Schaum, 1863, Cylindera [Cicindela] 111



foveolata Illiger, 1801, Sericoda [Carabus] 466

foveolatior Deuve, 1998, Carabus 187



foveolatoseriatus Reitter, 1896, Carabus 160

foveolatus Mulsant & Godart, 1860, Agabus 36

foveolatus M. F. Adams, 1817, Carabus 147

foveolatus Heer, 1839, Hydroporus 61

foveolatus Duftschmid, 1812, Pterostichus [Carabus] 506

foveolatus Gebler, 1841, Pterostichus [Platysma] [HN] 487

foveolatus Dejean, 1831, Syntomus [Dromius] 432

foveolatus Schaum, 1864, Zabrus 570

foveolus Rey, 1886, Notiophilus 97

foveostriatus C. G. Thomson, 1884, Haliplus 30

foveum Motschulsky, 1844, Bembidion 247

fowleri Deuve, 1993, Dongodytes 298

fowleri Omer‑Cooper, 1930, Gyrinus 28

fracassii Heyden, 1908, Calathus 524



fracta Fischer von Waldheim, 1828, Cicindela 107

fracta Motschulsky, 1844, Cicindela 106

fractilinea Solsky, 1874, Herophydrus [Hydroporus] 71

fractivittis Kraatz, 1890, Cicindela 105

fractus Sharp, 1882, Graptodytes [Hydroporus] 59

fragile Andrewes, 1925, Asaphidion 242

fragilis Uéno, 1989, Eocnides 308

fragilis Mateu, 1956, Microlestes 423

fragilis Mäklin, 1878, Pterostichus [Feronia] 490

franchettii Luigioni, 1926, Duvalius [Duvalites] 303



franciscoloi Sanfilippo, 1950, Duvalius 305

francki G. Ochs, 1929, Gyrinus 27

frankenbergeri Obenberger, 1914, Carabus 163

frankenbergeri Obenberger, 1913, Stomis 518

franki Roubal, 1929, Amara 551

franklinii Kirby, 1837, Dytiscus 52

franzi Hieke, 1981, Amara 563

franzi Jeannel, 1958, Apoduvalius 295

franzi Fassati, 1957, Bembidion 261

franzi Mandl, 1974, Carabus 162

franzi Wewalka & Biström, 1998, Glareadessus 55

franzi Mateu, 1954, Harpalus 376

franzi Zamotajlov & Sciaky, 1998, Ledouxius [DA] 284



franzi Machado, 1987, Licinopsis 540

franzi Dostal, 1986, Notiophilus 97

franzi Machado, 1992, Philorhizus 425

franzi Nègre, 1955, Pterostichus 503

franzi Dostal, 1993, Reicheiodes 230

franzi Schweiger, 1950, Trechus 329

franzianus Mateu & Deuve, 1979, Trechus 331

franziniorum Sciaky, 1987, Ophonus 391

frater Aubé, 1863, Anillus 238

frater Daffner, 1998, Anophthalmus 291



frater Kraatz, 1879, Carabus 199

frater Chaudoir, 1876, Chlaenius 349

frater Reitter, 1897, Leistus 82

frater Kunze, 1818, Nebrioporus [Hyphydrus] 66

frater Puel, 1938, Scarites 232

fraterculoides Breuning, 1961, Carabus 194

fraterculus Reitter, 1895, Carabus 149

fraterculus Maindron, 1899, Chlaenius 349



fraterna Putzeys, 1846, Aspidoglossa 219

fraternus Csiki, 1906, Carabus 172

fraudator Lapouge, 1905, Carabus 172

fraudulentus Spaeth, 1900, Notiophilus 97

fraxator Ménétriés, 1832, Bembidion 259

frederici J. Müller, 1918, Bembidion 267

fresnedai Fery, 1992, Hygrotus [Coelambus] 71

freudei V. B. Guéorguiev, 1975, Agabus 39

freudei Guignot, 1955, Copelatus 47

freudei Jedlika, 1966, Lebia 428

freudei Jedlika, 1965, Pterostichus 513

freudei Jedlika, 1965, Zabrus [Harpalus] 571

freyanus Uyttenboogaart, 1940, Poecilus [Pterostichus] 479

freyellus Jedlika, 1959, Bedeliolus 286

freyellus Jedlika, 1959, Chlaenius 349



freyellus Jedlika, 1958, Pterostichus 513

freyi Baliani, 1943, Amara 555

freyi Straneo, 1938, Aristochroa 471

freyi Netolitzky, 1937, Bembidion 259

freyi Colas, 1941, Calathus 526

freyi Jedlika, 1960, Calleida 408

freyi Emden, 1932, Carabus 176

freyi Jedlika, 1960, Chlaenius 351

freyi Schatzmayr, 1939, Cymindis 415

freyi Jedlika, 1958, Dyschirius 225

freyi Scheibel, 1935, Pterostichus 506



friaulicus Bernau, 1914, Carabus 179

friebi Jedlika, 1946, Amara 550

friebi Netolitzky, 1914, Bembidion 21, 250



friebianum Netolitzky, 1926, Bembidion 263

friedrichi Falkenström, 1936, Agabus [Gaurodytes] 39

friedrichi Falkenström, 1936, Rhantus 46

frigens Jeannel, 1928, Typhlotrechus 344

frigida Putzeys, 1867, Amara [Leirides] 563

frigida R. F. Sahlberg, 1844, Nebria 88



frigidum J. R. Sahlberg, 1880, Bembidion 265

frigidus Schiødte, 1841, Agabus 40

frigidus Fabricius, 1801, Calathus [Carabus] 525

frigidus Dejean, 1828, Pterostichus [Feronia] 492

frigidus J. Daniel, 1903, Pterostichus 503

frigidus Apfelbeck, 1904, Trechus 328

frigoris Csiki, 1930, Pterostichus 503

frioli Solier, 1835, Ditomus [Ditomus] 365

frischii Bergsträsser, 1778, Dytiscus 52

frischii Herbst, 1784, Pterostichus [Carabus] 509

frisium Paul Meyer, 1938, Bembidion 243

frisius Verhoeff, 1891, Hydroporus 64

fritzhiekei J. Schmidt, 1994, Pterostichus 510



friulanus J. Müller, 1926, Harpalus 373

frivaldskyi Kraatz, 1887, Carabus 170

frivaldszkyi Jakobson, 1906, Amara 551

frivaldszkyi Csiki, 1928, Bembidion 257

frivaldszkyi Roeschke, 1907, Cychrus 202

frivaldszkyi Reitter, 1893, Cymindis 415

frivaldszkyi Ganglbauer, 1903, Duvalius [Trechus] 299

frivola Bates, 1878, Amara [Leiocnemis] 564



froehlichii Duftschmid, 1812, Leistus [Carabus] 82

froelichii Sturm, 1818, Harpalus 376

frondicola Reitter, 1881, Reicheidius [Reicheia] 222

frontalis Sharp, 1882, Canthydrus 34

frontalis Putzeys, 1866, Dyschirius 226

frontalis Motschulsky, 1860, Dytiscus 52

frontalis Fischer von Waldheim, 1829, Harpalus 386



frontalis Sharp, 1882, Hyphydrus 74

frontalis Sharp, 1884, Leiodytes [Bidessus] 56



frontalis Hayward, 1900, Porotachys [Tachys] 274

frontalis Marsham, 1802, Rhantus [Dytiscus] 46

frontanyaensis Mollard, 1999, Carabus 141

frontosus Lapouge, 1905, Carabus 199

fruhstorfei W. Horn, 1902, Neocollyris [Collyris] 116



fruhstorferi Born, 1920, Carabus 184

fruhstorferi Roeschke, 1900, Carabus [Coptolabrus] 142



fruhstorferianus Csiki, 1927, Carabus 184

fuartensis Colla, 1993, Anophthalmus 292

fuchsi Scheibel, 1937, Duvalius 303

fuentei Breuning, 1926, Carabus [Mesocarabus] 167

fugatus Cavazzuti, 1998, Carabus 125

fugax Chaudoir, 1876, Chlaenius 349

fugax Faldermann, 1836, Harpalus 381

fugax A. Morawitz, 1862, Pterostichus 487

fugiens Andrewes, 1929, Oxylobus 231

fugitivus Chaudoir, 1868, Pterostichus [Feronia] 493

fujianensis Kleinfeld, 1997, Carabus 127

fujianensis Deuve & Tian, 1999, Leistus 80

fujianicus Deuve, 1998, Carabus 127

fujii Suenson, 1957, Epaphiopsis [Glabrotrechus] 309

fujiii Tanaka, 1959, Amara 556

fujimurai Habu, 1958, Pterostichus 493

fujisan Ishikawa, 1972, Carabus 149

fujisanus Bates, 1883, Carabus 159

fujisanus Uéno, 1971, Kurasawatrechus 313

fujisanus Tanaka & Suga, 1972, Pterostichus 501

fujitai Tanaka & Ishida, 1972, Pterostichus 498

fujitai Uéno, 1969, Trechiama 322

fujiwaraorum Uéno, 1981, Trechiama 322

fujiyamai Habu, 1958, Bembidion 265

fukiensis Jedlika, 1956, Amara 550

fukiensis Jedlika, 1956, Badister 441

fukiensis Jedlika, 1956, Dromius [Lebidromius] 420

fukiensis Jedlika, 1953, Eucolliuris 443

fukiensis Jedlika, 1956, Harpalus 386

fukiensis Jedlika, 1953, Lebia 428

fukiensis Jedlika, 1956, Lesticus 471

fukiensis Jedlika, 1956, Platynus [HN if Platynus] 464

fukiensis Jedlika, 1956, Platynus [Colpodes] 464

fukiensis Jedlika, 1957, Pseudognathaphanus [Kareya] 363

fukiensis Jedlika, 1953, Stenolophus [Egadroma] 405

fukiensis Jedlika, 1965, Tachyura [Tachys] 279

fukiensis Jedlika, 1956, Trichotichnus 395

fukuharai Habu, 1955, Synuchus [Calathus] 545

fukuharai Habu, 1957, Trichotichnus 395

fukukii Uéno, 1953, Epaphiopsis 309



fulgens Davis, 1832, Agonum 450

fulgens Charpentier, 1825, Carabus 165

fulgida Gebler, 1847, Nebria 89

fulgidicolle Erichson, 1841, Agonum [Anchomenus] 450

fulgidicollis L. Dufour, 1820, Chlaenius 352


fulgidus Fischer von Waldheim, 1828, Carabus 165

fulgidus Stephens, 1828, Chlaenius 350

fulgidus Motschulsky, 1850, Dyschirius 228



fulgidus Motschulsky, 1850, Poecilus 482

fuligineus A. Morawitz, 1862, Pterostichus 493

fuliginosa Dejean, 1826, Lophyra [Cicindela] 113

fuliginosum Panzer, 1809, Agonum [Carabus] 453

fuliginosum Netolitzky, 1914, Bembidion 259

fuliginosus Hölzel, 1950, Carabus 184

fuliginosus Duftschmid, 1812, Harpalus [Carabus] 382

fuliginosus Fabricius, 1792, Ilybius [Dytiscus] 42



fulminans Kraatz, 1878, Carabus [Sphodristus] 194

fulminifer G. Hauser, 1921, Carabus [Coptolabrus] 144

fulva Gené, 1839, Agelaea 449

fulva O. F. Müller, 1776, Amara [Carabus] 554

fulvaster Zaitzev, 1906, Agabus 37



fulvaster Motschulsky, 1861, Clivina [Eupalamus] 220

fulvatilis Belousov & Kabak, 1998, Trechus 331

fulveola Bates, 1883, Drypta 358

fulvescens Leach, 1819, Aepus 288

fulvescens Régimbart, 1889, Canthydrus 33

fulvescens Takizawa, 1931, Gyrinus 27

fulvescens A. Zimmermann, 1923, Hydroglyphus [Bidessus] 56

fulvescens Motschulsky, 1844, Pterostichus [Argutor] 490

fulvibasis Reitter, 1901, Microlestes [Blechrus] 422

fulvicollis Erichson, 1837, Haliplus 30

fulvicollis Fabricius, 1792, Lebia [Carabus] 427

fulvicollis Dejean, 1831, Tachys [Bembidium] 275

fulvicornis Bates, 1873, Stenolophus 405

fulvii Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 303

fulvilabris Dejean, 1826, Leistus 82

fulvipenne Schuler, 1959, Bembidion [Peryphus] 256

fulvipennis Régimbart, 1899, Agabus 36

fulvipennis Motschulsky, 1844, Harpalus 377

fulvipennis Munster, 1932, Hydroporus 64

fulvipennis Audinet‑Serville, 1821, Parophonus [Harpalus] 393

fulvipennis Dejean, 1831, Pogonus 287

fulvipennis Alluaud, 1939, Tachyura [Tachys] 277

fulvipes Audinet‑Serville, 1821, Amara [Harpalus] 567

fulvipes Sturm, 1827, Bembidion 249

fulvipes Gyllenhal, 1810, Calathus [Harpalus] 527

fulvipes Géhin, 1885, Carabus 136

fulvipes Latreille, 1817, Carterus [Aristus] 364



fulvipes Chaudoir, 1842, Chlaenius 349

fulvipes Motschulsky, 1839, Deltomerus [Platynus] 282



fulvipes Dejean, 1825, Dyschirius [Clivina] 228

fulvipes Della Beffa, 1909, Harpalus 375

fulvipes Fabricius, 1792, Harpalus [Carabus] 378

fulvipes Erichson, 1843, Idiomelas [Stenolophus] 404



fulvipes E. Jacquet, 1887, Lebia 428

fulvipes Motschulsky, 1845, Notiophilus 97

fulviventris Motschulsky, 1864, Brachinus 214

fulviventris A. Costa, 1883, Hygrotus [Hydroporus] 72

fulviventris Bassi, 1834, Nebria 92

fulvoides Jeanne, 1974, Trechus 23, 331

fulvoligatus Cavazzuti & Ratti, 1998, Carabus 156



fulvus Marsham, 1802, Bradycellus [Carabus] 401

fulvus Geoffroy, 1785, Demetrias [Buprestis] 419

fulvus Fabricius, 1801, Haliplus [Dytiscus] 32

fulvus Dejean, 1829, Harpalus 370

fulvus Chaudoir, 1846, Leistus 82

fulvus Alfieri, 1976, Microlestes 422

fulvus Baudi di Selve, 1864, Sirdenus [Pogonus] 288

fulvus Habu, 1978, Synuchus 545

fulvus Dejean, 1831, Trechus 331

fumatum Motschulsky, 1850, Bembidion [Philochthus] 264



fumatus Andrewes, 1924, Chlaenius 349

fumatus Sharp, 1882, Cybister 50

fumatus Schaum, 1854, Peronomerus 448

fumatus Mateu, 1961, Philorhizus 425

fumicata Motschulsky, 1851, Tachyura [Tachys] 277

fumidicolor Deuve, 1998, Carabus 174

fumigatoida Minowa, 1932, Tachyura [Tachys] [DA] 277

fumigatum Duftschmid, 1812, Bembidion [Elaphrus] 248

fumigatus Semenov, 1898, Carabus 189

fumipenne Fassati, 1957, Bembidion 262



funebris Sturm, 1827, Cicindela 103

funebris Schmidt‑Göbel, 1846, Cylindera [Cicindela] 109

funerarius Tschitschérine, 1890, Pterostichus [Feronia] 485

funerea W. S. MacLeay, 1825, Calomera [Cicindela] 100

funerea Ledoux & Roux, 1992, Nebria 94



funereum Ragusa, 1883, Calosoma 119

funerulum Fairmaire, 1886, Cymbionotum 234

funestes Csiki, 1930, Pterostichus 512

funestum Géhin, 1885, Calosoma 120

funestus Schilsky, 1888, Graptodytes [Hydroporus] 59

funestus Audinet‑Serville, 1821, Harpalus 380

funkii Seidlitz, 1887, Agabus 36

fur Andrewes, 1930, Platynus [Anchomenus] 462

furax Csiki, 1927, Carabus 187


furcatus Seidlitz, 1887, Haliplus 30

fureoiensis Kanô, 1922, Carabus 154

furihatai Morita, 1997, Trichotichnus 395

furumielloides Deuve, 1997, Carabus 122

furumiellus Deuve, 1994, Carabus 144



furumii Mandl, 1982, Carabus 122

furumii Deuve, 1990, Cychrus 204



furviculus Casey, 1924, Harpalus 377

furvus Tschitschérine, 1895, Calathus 527



furvus Guignot, 1950, Hydrovatus 70

furvus Machado, 1992, Paradromius 424


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