Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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alluaudi Antoine, 1957, Zabrus 572

almasyi Fassati, 1990, Bembidion 245

almeriensis Mateu, 1954, Acinopus 367

almeriensis Mateu, 1952, Tachyura 278

almonius Reitter, 1903, Trechus 326

almorae Andrewes, 1920, Chlaenius 353

almorensis Champion, 1923, Protopaussus 211

almosi Bokor, 1921, Duvalius [Trechus] 300

almum J. R. Sahlberg, 1900, Bembidion 247

almushensis Mandl, 1975, Carabus 194


alonsoi Cortés, 1988, Chlaenius 354

alotrus Lassalle & Remond, 1998, Carabus 154

alpestre Heer, 1838, Agonum [Anchomenus] 449



alpestris Heer, 1837, Agabus 38

alpestris A. Villa & G. B. Villa, 1833, Amara 563

alpestris Daffner, 1996, Anophthalmus 292

alpestris Sturm, 1815, Carabus 180



alpestris Beuthin, 1888, Cicindela 105

alpestris Heer, 1837, Cicindela 105

alpestris L. Redtenbacher, 1849, Harpalus 381

alpestris Falkenström, 1939, Hydroporus 60

alpestris Dejean, 1828, Molops [Feronia] 472



alpestris Gistel, 1857, Molops [HN] 472

alpestris Heer, 1837, Pterostichus [Argutor] 493

alpestris Heer, 1837, Trichotichnus [Harpalus] 396

alpherakii Semenov, 1898, Carabus 146

alphonsi J. Müller, 1914, Anophthalmus [Trechus] 290

alpicola Zaitzev, 1927, Agabus 39

alpicola Dejean, 1828, Amara 563



alpicola Heer, 1837, Amara 548

alpicola Jeannel, 1941, Bembidion [Peryphus] 269



alpicola Heer, 1837, Carabus 136

alpicola Fuss, 1860, Leistus 82

alpicola Motschulsky, 1866, Nebria [Alpaeus] 85

alpicola Mulsant & Rey, 1853, Pterostichus [Feronia] 512

alpicola Ganglbauer, 1891, Reicheiodes [Dyschirius] 230

alpicola Sturm, 1825, Trechus 326

alpigrada Csiki, 1906, Nebria 87

alpigradus Schauberger, 1927, Abax 470



alpigradus Sermet, 1985, Carabus 171

alpigradus Reitter, 1888, Trechus 326

alpina Paykull, 1790, Amara [Carabus] 558

alpina Gistel, 1837, Cicindela 105

alpina Chaudoir, 1873, Cymindis 411

alpina Heer, 1838, Loricera 98

alpinum Motschulsky, 1844, Agonum 454

alpinum Dejean, 1831, Bembidion 259

alpinum J. Jacquet, 1935, Bembidion [Testediolum] 269

alpinus Motschulsky, 1860, Agabus [Colymbetes] 35

alpinus Dejean, 1829, Aptinus 212

alpinus Dejean, 1828, Calathus 527

alpinus Dejean, 1826, Carabus 180

alpinus Kwon & Lee, 1984, Carabus [Lipaster] 174

alpinus Dejean, 1828, Laemostenus [Pristonychus] 536

alpinus Paykull, 1798, Oreodytes [Dytiscus] 67


alpinus Curtis, 1827, Patrobus [NO] 22, 285

alpinus Meschnigg, 1934, Philorhizus [Dromius] 425

alpinus Baudi di Selve, 1871, Scotodipnus 240

alpinus Heer, 1838, Synuchus 546

alpinus Dejean, 1831, Trechus 340

alpinus Morita, 1997, Trichotichnus 395

alpivagus Tschitschérine, 1899, Harpalus 372

alpujarrensis Mateu & Colas, 1954, Acupalpus 399

alsaticus E. Scriba, 1905, Haliplus 31

alstoni Péringuey, 1888, Brachinus [Pheropsophus] 212

altaianus Jedlika, 1958, Pterostichus 505



altaica Jedlika, 1957, Amara [Zabrus] 565

altaica Eschscholtz, 1829, Cicindela 103

altaica Gebler, 1833, Cymindis 416

altaica Jedlika, 1968, Cymindis 416

altaica Gebler, 1847, Nebria 94



altaica Jedlika, 1954, Rhopalostyla [Lebia] 430

altaicum Gebler, 1833, Bembidion 265



altaicus Motschulsky, 1844, Amara [Leirus] 559

altaicus Motschulsky, 1864, Brachinus 213

altaicus Gebler, 1847, Carabus 149

altaicus Jedlika, 1968, Harpalus 383

altaicus Gebler, 1848, Ilybius 43

altaicus Jedlika, 1968, Pseudotaphoxenus [Taphoxenus] 542

altaicus Germar, 1824, Pterostichus [Poecilus] 505

altaiensis Poppius, 1906, Pterostichus [Cryobius] 489

altamontanus J. Ochs, 1965, Carabus [Chrysocarabus] 140

altanbulagi Jedlika, 1968, Cymindis 417

alter Semenov & Znojko, 1929, Martyr 443



alternans Motschulsky, 1845, Bembidion [Notaphus] 254

alternans Faldermann, 1836, Calathus 524



alternans Kraatz, 1887, Carabus 179

alternans Palliardi, 1825, Carabus 161



alternans Chaudoir, 1856, Chlaenius 348

alternans Rambur, 1837, Cymindis 411



alternans Gravenhorst, 1807, Hygrotus [Dytiscus] 72

alternans Dejean, 1828, Licinopsis [Pristonychus] 540



alternans Carret, 1897, Pterostichus [Omaseus] 499

alternans Bedel, 1905, Singilis 438

alternans Balkenohl, 1999, Trilophus 222

alternatus Haury, 1885, Carabus 151

alternatus Blackburn, 1894, Celaenephes [Taromorpha] 410

alternatus Breit, 1911, Poecilus [Pterostichus] 484

altestriatum Netolitzky, 1934, Bembidion 243



alticola Bruneau de Miré & Legros, 1963, Agabus 39

alticola A. Fiori, 1903, Bembidion 260

alticola Britton, 1948, Bradycellus 400



alticola Bellier, 1880, Carabus 171

alticola Andrewes, 1926, Cymindis 412

alticola Bates, 1878, Dicheirotrichus 402

alticola Laneyrie, 1952, Duvalius 304

alticola Uéno, 1989, Epaphiopsis 309



alticola Jeannel, 1941, Geotrechus 309

alticola Sharp, 1882, Hydroporus 63

alticola Uéno & Saito, 1991, Jujiroa 459



alticola Leoni, 1907, Laemostenus 533

alticola Wollaston, 1864, Masoreus 357

alticola Bates, 1891, Morphodactyla [Pristodactyla] 530

alticola Grouvelle, 1913, Omoglymmius [Rhysodes] 78

alticola Jeannel, 1928, Orotrechus 317

alticola Wollaston, 1864, Platyderus 521

alticola Antoine, 1956, Pogonus 287

alticola Löbl, 1995, Sphaerius [Microsporus] 26



alticola Iablokoff‑Khnzorian, 1971, Trechus [Altaiotrechus] 334

alticola Wollaston, 1854, Trechus 326

alticola J. Schmidt, 1998, Xestagonum 467

alticola Fairmaire, 1889, Xestopus [Pristonychus] 530



altimirai Lagar Mascaró, 1964, Chlaenius 353

altiphila Hieke, 1995, Amara 552

altissima Hieke, 1981, Amara 552

altitudinis Ledoux & Roux, 1996, Archastes 79



altitudinorum Breuning, 1943, Carabus [Megodontus] 164

altitudinum Breuning, 1934, Carabus 164

altitudinum Deuve & Mourzine, 1998, Cychrus 205



altomontensis Branes, 1987, Cychrus 205

altyntuanus Shilenkov, 1996, Carabus 172

alutacea Lesne, 1896, Clivina 219

alutaceus Gebler, 1829, Chlaenius 348



alutaceus Sturm, 1826, Cychrus 203

alutaceus Wollaston, 1857, Dromius 419

alutaceus Tschitschérine, 1893, Poecilus [Feronia] 481

alutensis Savulescu, 1972, Carabus 172



alveolata Bates, 1892, Mastax 217

alveolatus Semenov, 1898, Carabus 145

alveolatus Chaudoir, 1879, Oxylobus 231



alyshensis Deuve & Quéinnec, 1992, Trechus 337

alysidotus Illiger, 1798, Carabus 191

alzonai Deuve, 1990, Carabus 156

alzonai Deuve & Ledoux, 1989, Nebria 90

alzonai Binaghi, 1935, Pterostichus 511


amabilis Hampe, 1871, Amara 567

amabilis Reitter, 1919, Brachinus 212

amabilis L. Redtenbacher, 1867, Callistomimus 348

amabilis Dejean, 1831, Cylindera [Cicindela] 110

amabilis Chaudoir, 1852, Drypta 358

amabilis Ledoux, Roux & Sawada, 1991, Nebria 88

amabilis Schaufuss, 1863, Neotrechus [Anophthalmus] 315



amabilis Tschitschérine, 1893, Poecilus [Feronia] 481

amaenus Chaudoir, 1859, Metacolpodes [Colpodes] 460

amagisanum Morita, 1994, Negreum [Colpodes] 460

amagisanus Tanaka & Ishida, 1972, Pterostichus 498

amamiensis Satô, 1961, Hydroglyphus [Guignotus] 56

amamiooshimensis Habu, 1964, Parena 433

amamioshimae Habu, 1978, Synuchus 544

amamioshimense Habu, 1955, Habragonum [Agonum] 458

amandatus Antoine, 1939, Leistus 84

amani Breit, 1933, Tapinopterus 519

amanoi Nakane, 1968, Pterostichus 497

amarae Andrewes, 1920, Chlaenius 353

amarellus Bates, 1891, Harpalus 372

amariformis Motschulsky, 1844, Harpalus 372

amariformis Bates, 1892, Prakasha [Platymetopus] 393

amaroides Reitter, 1890, Amara [Harpalodema] 547

amaroides Piochard de la Brûlerie, 1875, Amphimasoreus 521



amaroides Faldermann, 1836, Harpalus 383

amaroides Putzeys, 1877, Pristosia [Calathus] 530

amaroides Dejean, 1828, Pterostichus [Feronia] 489

amasiae Semenov, 1891, Laemostenus 537

amasicus Csiki, 1927, Carabus 185

amasiensis Reitter, 1900, Harpalus 380

amator Reitter, 1900, Harpalus 374

amaurocephalus Kolenati, 1845, Trechus 339

amaurum Bates, 1883, Bembidion 243

amauryi Haury, 1898, Carabus 201

ambicornis Sokolá, 1907, Carabus 196

ambiellensis Jeannel, 1948, Hydraphaenops [Aphoenops] 311



ambigena Bates, 1878, Amara 557

ambigenus Reiche, 1853, Harpalus 373

ambigenus Bates, 1883, Pterostichus 510



ambigenus Desbrochers des Loges, 1906, Pterostichus [HN] 489

ambigua Glasunov, 1902, Nebria 90



ambigua Mäklin, 1857, Sericoda [Anchomenus] 466

ambiguum Dejean, 1831, Bembidion 252

ambiguum Fairmaire, 1891, Xestagonum [Colpodes] 467

ambiguus Dejean, 1829, Acinopus 367



ambiguus Bates, 1883, Apatrobus [Penetretus] 280

ambiguus Paykull, 1790, Calathus [Carabus] 526



ambiguus Lapouge, 1914, Carabus 157

ambiguus Csiki, 1931, Chlaenius 352

ambiguus Aubé, 1838, Hydroporus 65

ambiguus Dejean, 1825, Pheropsophus [Brachinus] 217

ambiguus Fairmaire, 1858, Pterostichus [Feronia] 485

ambiguus Rambur, 1838, Zabrus 569

ambitiosus Schaufuss, 1872, Carabus 173

ambitus Laneyrie, 1948, Pterostichus [Troglorites] 514

amblygonellus Jeannel, 1964, Trechus 326

amblygonus Jeannel, 1926, Chaetoduvalius [Duvalius] 297

amblygonus Jeannel, 1960, Trechus 326

amblyops Bolívar y Pieltain, 1940, Pseudoplatyderus 524

amblypterus Chaudoir, 1868, Pterostichus [Feronia] 489

ambroggii Antoine, 1961, Laemostenus [Pristonychus] 533

ambrolauricus Belousov, 1989, Trechus 326

ambulans C. Zimmermann, 1832, Amara 555

ambulans Deuve, 1992, Cychrus 205

ambulator Andrewes, 1930, Xestagonum [Anchomenus] 467

amdoensis Hieke, 1996, Amara 552

amdoensis Semenov, 1889, Harpalus 377

amdoensis Deuve & Quéinnec, 1993, Trechus 333

americana Motschulsky, 1864, Diacheila 206

americanus Sharp, 1882, Hydaticus 54

amethystinum Meier, 1899, Bembidion 247

amethystinum Semenov & Redikorzev, 1928, Calosoma 118



amethystinus Dejean, 1828, Laemostenus [Pristonychus] 536

amicorum Wrase, 1993, Eocarterus 365

amicorum Zaballos, 1998, Geocharis 239

amicorum Uéno, 1999, Kusumia 314

amicorum J. Schmidt, 1996, Platynus 463

amicorum Jaeger, 1997, Psychristus 404

amicorum P. Moravec & Wrase, 1998, Trechus 326

amicta Wollaston, 1864, Cymindis [Tarus] 417



amiculus Guignot, 1948, Canthydrus 34

amicus Kryzhanovskij, 1995, Andrewesius 456



amicus L. Dufour, 1843, Harpalus 373

amicus Reitter, 1900, Harpalus 380

ammophila Tschitschérine, 1898, Bleusea 369

ammophilus Dejean, 1829, Acinopus 369

amnicola J. R. Sahlberg, 1880, Agabus [Gaurodytes] 37

amnicola J. R. Sahlberg, 1900, Bembidion 243

amoena Faldermann, 1835, Diacheila [Blethisa] 206



amoena Dejean, 1829, Dioryche [Platymetopus] 369

amoenulum Péringuey, 1896, Bembidion 254

amoenulus Semenov, 1889, Agatus [Glycia] 408

amoenulus Jedlika, 1934, Metacolpodes [Colpodes] 460

amoenulus Reitter, 1900, Ophonus 389

amoenum R. F. Sahlberg, 1844, Bembidion 266

amoenus Solsky, 1874, Agabus 35

amoenus Baudet‑Lafarge, 1836, Carabus 171

amoenus Chaudoir, 1852, Carabus 163

amoenus Chaudoir, 1861, Catascopus 435

amoenus Olivier, 1795, Haliplus [Dytiscus] 30

amoenus Heer, 1837, Harpalus 381

amoenus Fischer von Waldheim, 1844, Laemostenus [Pristonychus] 538

amoenus Wollaston, 1864, Paradromius [Dromius] 424

amoenus Dejean, 1828, Pterostichus [Feronia] 489

amoenus J. R. Sahlberg, 1908, Scarodytes [Hydroporus] 68

amorei Ganglbauer, 1909, Percus 478

amorei Ganglbauer, 1907, Pterostichus 503

amphibolus J. Müller, 1922, Dyschirius 227

amphinomum Bates, 1883, Nipponagonum [Colpodes] 460

ampliata Bates, 1873, Amara [Bradytus] 553

ampliatipennis Carret, 1904, Pterostichus 513

ampliatus Zaitzev, 1927, Hydroporus 60



ampliceps J. R. Sahlberg, 1921, Hydroporus 65

amplicollis Mannerheim, 1853, Amara [Celia] 556

amplicollis A. Villa & G. B. Villa, 1844, Carabus 180

amplicollis Bleuse, 1914, Carabus 168

amplicollis Belousov, 1999, Caucasorites 297

amplicollis Ménétriés, 1848, Harpalus [NP] 372

amplicollis Toledo, 1994, Hydronebrius 41



amplicollis Fairmaire, 1880, Leistus 82

amplicollis Fairmaire & Laboulbène, 1854, Pterostichus [Feronia] 514

amplicollis Fairmaire, 1859, Trechus 326

amplicollis Uéno, 1955, Trephionus 546

amplior Deuve, 1994, Carabus 121

amplior Machado, 1992, Paradromius 424

amplipennis Baliani, 1943, Amara 553

amplipennis Lapouge, 1925, Carabus 179

amplipennis Bates, 1873, Dicheirotrichus 403

amplipennis J. Müller, 1911, Neotrechus [Trechus] 315

amplipennis Roubal, 1928, Philorhizus [Dromius] 426

amplissimus O. F. Müller, 1776, Dytiscus 52

amplius Escalera, 1921, Calathus 526

amplum J. R. Sahlberg, 1908, Bembidion 20, 249

amplus Joseph, 1871, Anophthalmus 291



amplus J. Müller, 1916, Molops 472

amputatoides Mlyná, 1979, Harpalus 372

amrishi Makhan, 1999, Haliplus 32

amurense Motschulsky, 1859, Bembidion [Peryphus] 271



amurense Tschitschérine, 1895, Bembidion 250

amurensis Lutshnik, 1935, Amara 549

amurensis Géhin, 1885, Carabus 135

amurensis A. Morawitz, 1862, Cylindera [Cicindela] 110

amurensis Motschulsky, 1860, Demetrias 418

amurensis Fedorenko, 1991, Dyschirius 228



amurensis J. Balfour‑Browne, 1944, Dytiscus 52

amurensis Andrewes, 1924, Lebidia 426



amurensis Sharp, 1882, Nebrioporus [Deronectes] 65

amurensis Reitter, 1887, Philorhizus [Dromius] 426

amurensis Poppius, 1906, Pterostichus [Cryobius] 489



amurlandicus Shilenkov, 1994, Carabus 171

ana Uéno, 1955, Ja 458



anachoreta Ménétriés, 1832, Pterostichus [Feronia] 485

anadoluensis Kataev, 1993, Harpalus 381



anadyricus Csiki, 1930, Pterostichus 492

anae Morvan, 1982, Trechus 326

anagae Medina & Oromí, 1991, Orthomus [Wolltinerfia] 477

analis Fabricius, 1792, Amara [Carabus] 553

analis Aubé, 1838, Hydroporus 60

analis Jedlika, 1963, Pterostichus 514

analogicum Paul Meyer, 1919, Bembidion 255

anami Ledoux, 1977, Carabus 154

anamurensis Jedlika, 1965, Metadromius 421

ananowi Semenov, 1897, Carabus 169

anastomozans Well, 1781, Dytiscus 52

anatolica Motschulsky, 1859, Calomera [Cicindela] 99

anatolica Jedlika, 1965, Nebria 86

anatolica Jedlika, 1965, Tachyura [Tachys] [DA] 278



anatolicum Jedlika, 1968, Bembidion 252

anatolicum Korge, 1964, Bembidion 267

anatolicus Schweiger, 1968, Badister 440

anatolicus Wewalka, 1972, Bidessus 54

anatolicus Jedlika, 1969, Calathus 528

anatolicus Chaudoir, 1857, Carabus [Procrustes] 186



anatolicus Jedlika, 1962, Carterus 363

anatolicus Motschulsky, 1865, Cychrus 202



anatolicus K. Daniel, 1904, Harpalus [Ophonus] 374

anatolicus Tschitschérine, 1898, Harpalus 372

anatolicus J. Balfour‑Browne, 1963, Hydroporus 60

anatolicus Guignot, 1957, Hyphydrus 74

anatolicus Casale, 1988, Laemostenus 533


anatolicus Mlyná, 1977, Molops 473

anatolicus Coiffait, 1956, Parvocaecus [Winklerites] 240

anatolicus Jedlika, 1968, Parvocaecus [Scotodipnus] [DA] [HN!] 240

anatolicus Jedlika, 1963, Platyderus 522

anatolicus J. Schmidt, 1996, Platynus 462

anatolicus Chaudoir, 1850, Poecilus [Feronia] 481

anatolicus Jedlika, 1963, Pterostichus 489

anatolicus Ganglbauer, 1900, Trechus 336

anatolicus Ganglbauer, 1915, Zabrus 572

anaulax Uéno, 1978, Yamautidius 345

anaunus Depoli, 1939, Abax 470

anceps Csiki, 1927, Carabus 143

anceps Lapouge, 1925, Carabus 130

anceps Chaudoir, 1876, Chlaenius 350

anceps Putzeys, 1861, Clivina 220

anceps Putzeys, 1875, Tachyura [Tachys] 278

ancestralis Belousov, 1998, Nannotrechus 314

anchocephalus Reitter, 1896, Carabus 147

anchomenoides Bates, 1873, Bradycellus [Tachycellus] 401

anchomenoides Bates, 1889, Chlaenius 353

anchoralis Chevrolat, 1845, Abroscelis [Cicindela] 99

anchoralis Sharp, 1884, Nebrioporus [Deronectes] 66

ancora Ménétriés, 1832, Badister 440

ancorifer Uéno, 1995, Trechiama 321

andalusia Rambur, 1837, Nebria 92

andalusiacus Rambur, 1837, Brachinus 212



andalusiacus Géhin, 1885, Carabus 161

andalusiacus Jedlika, 1950, Chlaenius 352

andalusiacus J. Vives & E. Vives, 1982, Laemostenus [Ceuthosthenes] 533

andalusiacus Mateu, 1957, Orthomus 477

andalusiacus Jeannel, 1927, Trechus 331

andalusiae Clark, 1862, Nebrioporus [Hydroporus] 66

andalusicus Wehncke, 1872, Haliplus 31

andalusicus Reitter, 1897, Penetretus [Deltomerus] 285

andalusicus Motschulsky, 1862, Polyderis 274

andalusius Rambur, 1837, Mesolestes [Dromius] 421

andamanensis Pouillaude, 1914, Hexagonia 407

andamanicus Régimbart, 1899, Copelatus 47

andarensis Bolívar y Pieltain, 1923, Nebria 85

anderschii Duftschmid, 1812, Pangus [Carabus] 392

andiaensis Forel & Leplat, 1998, Carabus 179

andoi Uéno, 1959, Rakantrechus 319



andorranus Barthe, 1909, Carabus 168

andorranus Español & Comas, 1984, Hypotyphlus 239

andorranus Jeannel, 1937, Pterostichus [Haptoderus] [HN!] 489


andorranus Puel, 1924, Pterostichus 498

andorranus Jeanne, 1972, Trechus 339

andreae Tschitschérine, 1898, Amara 559

andreae Ahrens, 1812, Bembidion 243


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