Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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kaynasliensis P. Machard, 1975, Carabus 157

kazakhstanicum Kryzhanovskij, 1979, Bembidion 243

kazanensis Jedlika, 1958, Harpalus 378

kazanensis Belousov & Kabak, 1992, Trechus 330

kaznakovi Tschitschérine, 1903, Aristochroa 471

kaznakovi Semenov & Znojko, 1932, Carabus 150

kaznakovi Semenov & Znojko, 1934, Cychrus 204

kaznakovi Lutshnik, 1915, Cylindera [Cicindela] 110

kaznakovi Kataev & Wrase, 1997, Harpalus 378

kaznakovi Jeannel, 1935, Trechus 333

kazuyoshii Morita & Kurosa, 1994, Platynus 463

kazuyukii N. Ito, 1999, Trichotichnus 394

kebekensis Zamotajlov, 1988, Deltomerus 282



keeni Casey, 1918, Amara 549

keilbachi Müller‑Motzfeld, 1978, Bembidion 246

keinigus Morvan, 1994, Synuchus 545

keithi Deuve, 1995, Carabus 125

keithi Deuve, 1998, Cychrus 204

kekesiensis Nyilas, 1993, Poecilus 479

kellneri Hubenthal, 1910, Carabus 171

keltepensis Vigna‑Taglianti & Bruschi, 1976, Carabus 193

keltiekus Morvan, 1981, Pterostichus 510

keminensis Kabak, 1994, Carabus 145

kemm Morvan, 1999, Xestagonum 468

kempi Andrewes, 1922, Bembidion 258

kempi Gschwendtner, 1936, Hyphoporus 73


kempi Gschwendtner, 1936, Laccophilus 76

kempi G. Ochs, 1925, Orectochilus 29

kempi Vazirani, 1970, Platambus [Agraphis] 43

kenaiensis Fall, 1926, Ilybius [Agabus] 43

kenderessyi Mallász, 1924, Duvalius [Anophthalmus] 298

kendii Csiki, 1923, Carabus 179

kenteiana Lutshnik, 1935, Amara 550

kenyeriae Mandl, 1983, Chlaenius 349



kenyeryae Kenyery, 1977, Carabus 155

kenyeryi Schweiger, 1965, Carabus 181

kenzanensis Ishida & Shibata, 1961, Amara [Armatoleirides] 552

kerberos Kirschenhofer, 1997, Tapinopterus 520

kergreni Morvan, 1999, Xestagonum 468



keridwen Morvan, 1995, Xestagonum [Colpodes] 468

kermanensis J. Balfour‑Browne, 1939, Ilybiosoma [Agabus] 41

kerteszi Csiki, 1912, Anophthalmus 291

kervillei Puel, 1938, Calathus 526

kerzhneri Shilenkov, 1982, Nebria 88

kerzhneri Lafer, 1983, Pterostichus 486

kesharishresthae Morvan, 1999, Agonum 452



kessleri Hochhuth, 1871, Agabus 37

ketmenensis Dolin, 1991, Carabus 145

ketmenicus Shilenkov & Kabak, 1994, Leistus 82

ketmenicus Belousov & Kabak, 1993, Trechus 333

kevnidennek Morvan, 1998, Bruskespar 457

kezadonicus Belousov, 1989, Trechus 333

kezukai Deuve & Imura, 1991, Carabus 188

kezukai Dostal, 1984, Chaetobroscus 236

khalabicus Belousov, 1990, Trechus 333

khalafi Ali, 1967, Distichus [Scarites] 231

khaledicus Belousov, 1990, Trechus 333

khalisensis Ali, 1967, Acinopus 369

khan Casale, 1988, Pseudotaphoxenus 541

khanakense Mikhailov, 1984, Bembidion 256

kharazii Morvan, 1975, Brachinus 216

kharazii Morvan, 1974, Nebria 91

kharoumensis Mateu, 1981, Microdaccus 422

khersonensis Motschulsky, 1845, Cicindela 107

khnzoriani Kurnakov, 1960, Deltomerus 282

khnzoriani Kalashyan, 1983, Laemostenus [Pristonychus] 538

khnzoriani Pawowski, 1976, Trechus 333

kholini Nilsson, 1994, Agabus 40

khorasanensis Deuve, 1993, Carabus 169

khorassanica Ali, 1978, Calomera [Cicindela] 100

khorgosicus Belousov & Kabak, 1994, Trechus 333



khotanensis Deuve, 1992, Carabus 160

khukri Brancucci, 1990, Platambus 43



khyber Andrewes, 1935, Bembidion 254

kiangsianus G. Hauser, 1912, Carabus [Coptolabrus] 141

kiangsu Jedlika, 1965, Pterostichus 515

kiapazicus Belousov, 1990, Trechus 333

kiautschouensis Schauberger, 1930, Harpalus 385

kidanicus Kataev, 1989, Harpalus 376



kienchanganus G. Hauser, 1926, Carabus [Coptolabrus] 143

kienchangensis G. Hauser, 1918, Carabus [Apotomopterus] 124



kiesenwetteri Kraatz, 1872, Ilybius 41

kiesenwetteri Wehncke, 1872, Ilybius 42

kiesenwetteri Chaudoir, 1868, Pterostichus [Feronia] 490

kiesenwetteri Pandellé, 1867, Trechus 326

kiesenwetteri Schaum, 1862, Typhlotrechus [Anophthalmus] 344

kiesenwetterii Seidlitz, 1887, Agabus 39

kifunei Nakane, 1987, Hydroglyphus [Guignotus] 56

kiiensis Nakane & Iga, 1953, Carabus [Apotomopterus] 178

kiiensis Morita & Ohkura, 1988, Pterostichus 507

kikuyai Uéno, 1987, Rakantrechus 319

kildroennus Morvan, 1995, Platynus [Colpodes] 465

kiliani Peyerimhoff, 1929, Nebrioporus [Hydroporus] 67

kilikiensis Mandl, 1961, Homodela [Cicindela] 112

kimak Belousov & Kabak, 1996, Trechus 333

kimakowiczi Ganglbauer, 1891, Duvalius [Trechus] 307

kimieae Kezuka & Imura, 1989, Carabus [Damaster] 122

kimunkamui Nakajima, 1993, Carabus [Acoptolabrus] 122

kimurai Ishikawa, 1966, Carabus [Apotomopterus] 177

kimurai Morita, 1994, Pterostichus 513

kimurai Uéno, 1959, Trechiama 322



kindermanni Chaudoir, 1850, Carabus 192

kindermanni Hampe, 1852, Carabus 153

kindermanni Waltl, 1838, Carabus 187

kindermanni Chaudoir, 1856, Chlaenius 350

kindermanni Chaudoir, 1871, Graphipterus 359

kindermanni Chaudoir, 1861, Siagona 218



kineli Makolski, 1952, Badister 440

kinfushanicus Schauberger, 1933, Harpalus 384

kinfushanum Jedlika, 1958, Bembidion 255

kingdoni Baliani, 1934, Amara 549

kinitzi Tschitschérine, 1899, Amara 547

kinoshitai Habu, 1954, Trephionus 546

kintrishensis Gottwald, 1980, Carabus 201

kinzelbachi Fery & Hosseinie, 1998, Deronectes 58

kionchochi Zamotajlov, 1996, Deltomerus 282


kirbii Marsham 1802, Gyrinus 27

kirbyi Fischer von Waldheim, 1828, Carabus 135

kirbyi Schmidt‑Göbel, 1846, Hexagonia 407

kircheri Germar, 1838, Carabus 184



kirghisiaca Schilder, 1911, Cicindela 104

kirghisica Kryzhanovskij, 1962, Amara 566

kirghisorum Kryzhanovskij, 1953, Carabus 146

kirghizicus Fedorenko, 1994, Dyschirius 228

kirgisica Mandl, 1936, Cicindela 106

kirgisica Chaudoir, 1875, Cymindis 416

kirgisica Shilenkov, 1982, Nebria 90

kirgisicus Motschulsky, 1844, Harpalus 378

kirgisicus Komarov & Kabak, 1995, Philorhizus 425

kirgisiensis Mandl, 1967, Carabus 160

kirgisorum Netolitzky, 1934, Bembidion 268

kirikensis Staven, 1995, Carabus 194

kirilovii Fischer von Waldheim, 1844, Cylindera [Cicindela] 109

kirinicus Csiki, 1927, Carabus 156

kirishimanus Habu, 1973, Chydaeus 362

kirishimanus Habu, 1954, Pterostichus 487

kiritschenkoi Mikhailov, 1984, Bembidion 256



kiritschenkoi Breuning, 1934, Carabus 145

kiritshenkoi Kryzhanovskij, 1974, Amara [Curtonotus] 560

kiritshenkoi Emetz & Kryzhanovskij, 1973, Cymindis [DA] 415

kiritshenkoi Kataev, 1990, Harpalus 378

kiritshenkoi Lutshnik, 1928, Pterostichus [Platysma] 500

kirschenblatti Lutshnik, 1938, Bembidion 264

kirschenblatti Kryzhanovskij, 1988, Pterostichus 504

kirschenhoferi Wrase, 1985, Abax 470

kirschenhoferi Morvan, 1999, Aparupa 456

kirschenhoferi Müller‑Motzfeld, 1988, Bembidion [Hoquedela] 250

kirschenhoferi F. Battoni, 1982, Calathus 525

kirschenhoferi Morvan, 1992, Deltocolpodes 457

kirschenhoferi Straneo, 1983, Euryaptus 471

kirschenhoferi Casale, 1988, Laemostenus 534

kirschenhoferi Dostal, 1981, Notiophilus 97

kirschenhoferi Sciaky, 1994, Straneostichus 518

kirschenhoferianus F. Battoni, 1986, Calathus 525

kiselevi Kataev & Shilenkov, 1990, Harpalus 384

kishengangai Heinz, 1993, Carabus 155

kishii Nakane, 1963, Acilius 48

kishimotoi Morita, 1996, Bembidion [Ocydromus] 265

kishimotoi Uéno, 1998, Oodinotrechus 316

kishimotoi Morita, 1997, Trichotichnus 396



kiskunensis Ádám & Merkl, 1986, Carabus 197

kisonis Kasahara, 1995, Trichotichnus 396

kitaiensis Mandl, 1978, Pristomachaerus [Psuedochlaeniellus] 356

kitakamisanus Ishikawa, 1992, Carabus [Leptocarabus] 159

kitanoi Habu, 1956, Agonum 454

kitawakianus Imura, 1993, Carabus 190

kitawakiellus Imura, 1995, Carabus 189

kiuchii Uéno, 1969, Himiseus [Ryugadous] 311

kiukiangensis Bates, 1888, Carabus 156

kiushuensis Mandl, 1981, Chaetodera [Cicindela] 103

kizbaiensis Kabak, 1994, Poecilus 480

kjellanderi Luna de Carvalho, 1965, Paussus 210

klaegeri Kraatz, 1895, Carabus [Tropidocarabus] 155

klapperichi Jedlika, 1956, Amara 560

klapperichi Jedlika, 1953, Bembidion 271

klapperichi Jedlika, 1956, Brachinus 215

klapperichi Jaeger & Wrase, 1994, Bradycellus 401

klapperichi Jedlika, 1963, Calleida 408

klapperichi Mandl, 1942, Calomera [Cicindela] 99

klapperichi Mandl, 1955, Calosoma 119

klapperichi Reichensperger, 1954, Ceratoderus 209

klapperichi Jedlika, 1956, Chlaenius 356

klapperichi Jedlika, 1956, Cymindis 413

klapperichi Jedlika, 1964, Dixus [Ditomus] 365

klapperichi Jedlika, 1956, Glycia 409

klapperichi Jedlika, 1956, Harpalus 378

klapperichi Jedlika, 1953, Hexagonia 407

klapperichi Jedlika, 1953, Lebia 429

klapperichi Jedlika, 1956, Mastax 217

klapperichi Jedlika, 1953, Mochtheroides 437

klapperichi Bänninger, 1956, Nebria 90

klapperichi G. Ochs, 1942, Orectochilus 29

klapperichi Jedlika, 1956, Pterostichus 486

klapperichi Bänninger, 1956, Scarites 233

klapperichi Jedlika, 1953, Tachyura [Tachys] 279

klapperichi Mandl, 1955, Therates 116

klapperichi Jedlika, 1953, Trichotichnus [Asmerynx] 396

klapperichi Jedlika, 1956, Zabrus 572

klapperichianum Fassati, 1957, Bembidion 261

klapperichianus Mandl, 1955, Carabus 148

klarae Farka, 1995, Leistus 80

kleinfeldi Korell, 1980, Carabus 162

kleinfeldi Straneo, 1982, Pterostichus 510

kleinfeldianus Deuve, 1991, Carabus 127

kleinfeldianus Sciaky & Wrase, 1997, Pterostichus 501

kleinfeldorum Kabak & Putchkov, 1995, Carabus 159


kleini Schuler, 1964, Amara [Celia] 555

klementzae Kataev, 1984, Harpalus 381



klettei Gerhardt, 1899, Stenolophus 405

klickai Jedlika, 1928, Amara 558

klickai Jedlika, 1931, Chlaenius 349

klickai Jedlika, 1933, Lebia 429

klickai Jedlika, 1931, Platynus 464

klickai Jedlika, 1934, Platynus [Colpodes] [HN if Platynus] 465



klickai Jedlika, 1931, Pterostichus 497

klimenkoi Obydov, 1999, Carabus 164

klimeschi Winkler, 1914, Duvalius [Trechus] 307



klinckowstroemi Mjöberg, 1915, Nebria 93

klisuranus Apfelbeck, 1902, Molops 474

kluchoriensis Vysoký, 1981, Pterostichus 496

klugi Mannerheim, 1849, Carabus 149

klugi Nietner, 1858, Tachyura [Bembidium] 278



klugii Leprieur, 1880, Hydroglyphus [Hydroporus] 56

klugii Dejean, 1828, Pterostichus [Feronia] 488

klynstrai J. Müller, 1934, Dyschirius 227

klynstrai Andrewes, 1929, Rhysopus 363

klytchensis Gottwald, 1985, Carabus 132

knabli Mandl, 1951, Carabus 136

knauthi Ganglbauer, 1904, Boldoriella [Trechus] 296

knauthi Ganglbauer, 1901, Trichotichnus 396

kniephofi Langenhan, 1913, Carabus 195

knirschi Winkler, 1912, Anophthalmus [Trechus] 292

knirschi Jedlika, 1931, Brachinus 215

knirschi Janák & P. Moravec, 1989, Duvalius 299

knirschianus Daffner, 1996, Anophthalmus 291

kobachidzei Kurnakov, 1970, Carabus 131

kobayashie Habu, 1957, Trichotichnus 396

kobayashii Uéno, 1956, Suzuka [Ishidatrechus] 321

kobensis Sharp, 1873, Laccophilus 76

kobensis Habu, 1955, Synuchus [Calathus] 545



kobingeri Apfelbeck, 1904, Carabus 163

kobingeri Apfelbeck, 1902, Trechus 333



kocae Reitter, 1895, Bembidion [Limnaeum] 267

kocae Born, 1910, Carabus 195

kocae Ganglbauer, 1904, Laccornis [Hydroporus] 75

kocheri Antoine, 1933, Amara 555

kocheri Antoine, 1939, Cymindis 413

kocheri Verdier, 1951, Nebria 92

kocheri Mateu, 1955, Orthomus 477

kocheri Antoine, 1963, Paradromius 424

kocheri Antoine, 1956, Tachyura 278


kocheri Paulian & Villiers, 1939, Trechus 333

kochi Baliani, 1940, Amara 560

kochi Schatzmayr, 1937, Calathus 525

kochi Schatzmayr, 1936, Clivina 219

kochi Schatzmayr, 1944, Cymindis 414

kochi Schatzmayr, 1940, Nebria 93

kochi Schatzmayr, 1936, Zuphium 574

kociani Farka, 1999, Leistus 80

kocmani Kaláb, 1997, Carabus 175

kodarensis Brinev, 1997, Carabus 134

kodaricola Deuve, 1998, Carabus 134

kodoricus Belousov, 1989, Trechus 333

kodoricus Belousov, 1998, Troglocimmerites 344

kodoriensis Fedorenko, 1996, Reicheiodes 230



kodorii Morvan, 1970, Carabus 181

kodrici Scheibel, 1938, Duvalius [Neoduvalius] 307



kodymi Jedlika, 1936, Amara 549

kodymi Maan, 1939, Molops 474

kodymi Maan, 1940, Zabrus 570

koenigi Ganglbauer, 1887, Carabus 181

koenigi Semenov, 1888, Chlaenius 350

koenigi Zaitzev, 1927, Colymbetes 45

koenigi Reitter, 1887, Laemostenus [Antisphodrus] 533

koenigi Reitter, 1887, Philorhizus [Dromius] 425

koenigi Reitter, 1887, Pterostichus [Lyrothorax] 504

koenigianus Tschitschérine, 1890, Pterostichus [Feronia] 485

koeppeni Motschulsky, 1865, Laemostenus [Cryptoxenus] 538

kofleri Daffner, 1996, Anophthalmus 291

koganae Colas, 1961, Carabus 162

koheii Nakane, 1963, Pterostichus 498



kohistanicus Kirschenhofer, 1992, Harpalus 379

koidei Imura, 1996, Carabus 128

koikei Habu & Baba, 1960, Bembidion 271

koiwayai Hieke, 1993, Amara 560

koiwayai Zamotajlov & Sawada, 1996, Apatrobus 281

koiwayai Deuve & Imura, 1990, Carabus 125

koiwayai Deuve & Imura, 1993, Cychrus 204

koiwayai Ledoux & Roux, 1989, Nebria 91

koizumii Habu, 1961, Tachys [Eotachys] 275

kojedoensis Kwon & Lee, 1984, Carabus 183

kojensis Kurosawa & Kudo, 1980, Carabus [Damaster] 142

kokandicum Solsky, 1874, Bembidion 21, 269

kokeilii L. Miller, 1850, Pterostichus 506



kokensis Breuning, 1932, Carabus 145

kokoosson Feng, 1936, Agabus 36



kokshaalensis Deuve, 1993, Carabus 150

kokujewi Semenov, 1898, Carabus 138



kokujewi A. I. Jakovlev, 1896, Colymbetes [Cymatopterus] 45

kokujewi Tschitschérine, 1898, Thalassophilus [Trechus] 346

kokzhotensis Belousov & Kabak, 1996, Trechus 333



kolahoiensis Ledoux, 1980, Carabus 155

kolatensis Lapouge, 1925, Carabus 193

kolbei Roeschke, 1897, Carabus 164



kolbi Breuning, 1927, Carabus 181

kolbi Coiffait, 1972, Laemostenus [Pristonychus] 535



kolenatii Chaudoir, 1850, Asaphidion [Tachypus] 20, 242

kolenatii Chaudoir, 1846, Carabus 201



kolenatii Lutshnik, 1922, Harpalus 374

kollari Dejean, 1831, Bembidion 266

kollari Putzeys, 1873, Calathus 527

kollari Palliardi, 1825, Carabus 173

kollarisans Sokolá, 1907, Carabus 173

kolosovi Zinov'ev, 1997, Carabus 199

kolosovi Lutshnik, 1928, Pterostichus [Platysma] 515

kolshengelicum Obydov, 1997, Calosoma [Callisthenes] 118



kolstromi J. R. Sahlberg, 1875, Hydroporus 62

kolstromii C. R. Sahlberg, 1827, Bembidion 266

koltzei Paul Meyer, 1919, Bembidion 243

koltzei Reitter, 1900, Bradycellus 401

koltzei Rost, 1889, Carabus [Plectes] 131

koltzei Roeschke, 1907, Cychrus 205

kolymensis Lafer, 1989, Carabus 134

kolymensis Erjiomin, 1998, Pterostichus 491

kolyvanus Shilenkov & O. Berlov, 1996, Carabus 170

komala Jedlika, 1934, Amara 554

komala Andrewes, 1932, Platynus [Colpodes] 465

komalus Jedlika, 1936, Pterostichus [Neohaptoderus] 501

komareki Fassati, 1955, Bembidion 265

komareki Hrka & Pulpán, 1992, Nebria 87

komareki Jedlika, 1931, Pristosia [Kanoldia] 531

komarovi Zamotajlov, 1988, Deltomerus 282

komarovi Semenov & Znojko, 1928, Nebria 89

komarovi Lafer, 1976, Platynus [Agonum] 463

komarovi Belousov, 1990, Trechus 333

komarowi Reitter, 1882, Carabus 131

komarowi Semenov, 1889, Daptus 369

komarowi Semenov, 1889, Discoptera 356



komensis Horvatovich, 1973, Nebria 87

kominecki Bielz, 1851, Daptus 369

komiyai Ishikawa, 1966, Carabus [Apotomopterus] 177



kompeskanus Morvan, 1995, Platynus [Colpodes] 465

kondarensis Kryzhanovskij, 1953, Carabus 134



konduensis Gschwendtner, 1927, Cybister 50

konevi Kryzhanovskij, 1990, Bedeliolus 286

kongosanus Nakane, 1963, Pterostichus 502

koniensis Schauberger, 1927, Ophonus [Harpalus] 388

konkerianus Breuning, 1969, Carabus 163

konoi Uéno, 1955, Asaphidion 243

konoi Gotô & Iga, 1950, Cicindela 108

konoi Nakane, 1963, Hydroporus 65

konsenensis Ishikawa, 1968, Carabus [Damaster] 122

kopetdaghi Kryzhanovskij, 1968, Amara 566

koprivnicensis P. Moravec & Lompe, nom. nov., Trechus 22, 338

kopylensis Belousov & Kabak, 1994, Trechus 331

korabensis J. Müller, 1936, Deltomerus 283

korabensis Horvatovich, 1975, Nebria 86

korabica Horvatovich, 1974, Nebria 84

koralpicus Sokolá, 1910, Carabus 184

koralpicus Franz, 1969, Trechus 326

korax Reitter, 1889, Calathus 525



korbeli Majzlan, 1992, Duvalius 307

korbi Breuning, 1928, Carabus 186



korbi Beuthin, 1888, Cicindela 106

korbi Ganglbauer, 1900, Geocharis [Anillus] 239

korbi Ganglbauer, 1887, Omphreus 446

korbi Schauberger, 1933, Ophonus [Harpalus] 388

korbi Tschitschérine, 1893, Poecilus [Feronia] 482

korbi Reitter, 1903, Trechus 333

korbianus Ganglbauer, 1887, Carabus [Sphodristocarabus] 194

koreana Kirschenhofer, 1994, Parena 434

koreanica Mandl, 1981, Cylindera [Cicindela] 110

koreanorum Pawowski, 1974, Tachys 275

koreanus Reitter, 1895, Carabus 159

koreanus Mori, 1932, Copelatus 47



koreanus Takizawa, 1931, Peltodytes 33

koreanus Nilsson, 1997, Platambus [Agabus] 43



korelli Kleinfeld, 1988, Carabus 146

korelli Kleinfeld, 1999, Cychropsis 202



korgei Jedlika, 1964, Bradycellus [Stenolophus] 401

korgei Heinz, 1970, Carabus 192

korgei Breuning, 1964, Cychrus 202

korgei Jedlika, 1965, Nebria 86

korgei Jedlika, 1964, Pterostichus 491

korgei Jedlika, 1964, Styphlomerus [Brachinus] 218



korgei Pawowski, 1976, Trechus 335

korlevici Major Hoffmann, 1883, Carabus 151

kornreiza Morvan, 1995, Skoeda 466

korobeinikovi Shilenkov, 1996, Carabus 133

korolkowi Semenov, 1897, Carabus 192

korolkowi Tschitschérine, 1901, Pterostichus [Platysma] 502


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