Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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koropeckyi Deuve & Macek, 1992, Carabus 200

korotyaevi Deuve, 1991, Cychrus 204

korotyaevi Shilenkov, 1982, Trechus 333

korr Morvan, 1999, Xestagonum 468

korrigani Morvan, 1981, Deltomerus 282

korrigani Morvan, 1982, Trechus 333

korsakowi Semenov, 1898, Carabus 190

korschefskyi Eidam, 1931, Carabus [Diocarabus] 192

korzhun Belousov & Kabak, 1994, Trechus 333

kosagatschi Hieke, 1988, Amara 558

kosak Iljin, 1923, Brachinus 213

kosakai Morita, 1998, Pterostichus 498

kosakai Morita, 1997, Trichotichnus 396

koschagachensis Matalin, 1999, Cicindela 103



koselerensis P. Machard, 1991, Carabus 193

kosempoides Jedlika, 1952, Platynus [Colpodes] 465

kosensis Cavazzuti, 1984, Carabus 194

koshanini Csiki, 1904, Carabus 173

koshiensis Kirschenhofer, 1992, Harpalus 386

koshikicola Imura & Mizusawa, 1995, Carabus 177

koshtanensis Belousov & Zamotajlov, 1993, Carabus 200

kosti Matits, 1912, Bembidion 267

kosugei Nakane, 1955, Carabus [Procrustes] 164

kosugei Uéno, 1955, Trechiama 322

kotelensis Genest, 1978, Duvalius 308

kotoshonis Kanô & Kamiya, 1931, Haliplus 32

kotoshonis Kanô, 1931, Therates 115



kotschyi Letzner, 1849, Agabus 40

kotschyi Ganglbauer, 1887, Carabus [Procrustes] 188

kotschyi L. Redtenbacher, 1850, Eurynebria [Nebria] 79

kotulae Ulanowski, 1884, Acilius 48

kouanping Maindron, 1906, Carabus 125



kouanti Maindron, 1906, Carabus 128

koubbii Deuve, 1998, Carabus 157

kouraensis Puel, 1937, Dyschirius 224

kourili Maan, 1942, Molops 474

kourili Maan, 1933, Pterostichus 485



koutchii Ledoux, 1985, Nebria 90

kovali Belousov, 1999, Caucasorites 297

kovali Zamotajlov, 1988, Deltomerus 282


kovali Belousov, 1989, Nannotrechus 315

kovali Belousov, 1989, Trechus 333

koyi Germar, 1824, Poecilus 482

koyunensis Darge, 1990, Carabus 157

kozakai Perrault, 1984, Leistus 83

kozhantschikovi Lutshnik, 1924, Carabus 171

kozhantshikovi Lutshnik, 1924, Cicindela 105



kozhevnikovianus Lutshnik, 1933, Poecilus [Platysma] 480

koziorowiczi Piochard de la Brûlerie, 1873, Leistus 84

kozlovi Kryzhanovskij, 1995, Broscus 236

kozlovi Tschitschérine, 1903, Cicindela 104

kozlovi Semenov & Znojko, 1934, Cychrus 204

kozlovi Kabak, 1999, Cymindis 418

kozlovi Kataev, 1993, Harpalus 378



kozlovi Glasunov, 1902, Nebria 91

kozlovi Tschitschérine, 1903, Poecilus [Platysma] 480

kozlovi Jeannel, 1935, Trechus 333

kozloviellus Semenov & Znojko, 1932, Carabus 150

kozlovskii Zaitzev, 1927, Hydroporus 62

kozlowi Semenov, 1898, Carabus 178

kozlowi Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403

kraatzi Chaudoir, 1877, Carabus 200

kraatzi C. Müller, 1889, Oreonebria [Nebria] 96

kraatzi W. Horn, 1899, Pronyssa [Heptodonta] 115

kraatzi Heyden, 1882, Pseudotaphoxenus 541

kraatzi Andújar & Serrano, 2000, Zabrus 571

kraatzii Schaum, 1868, Hydroporus 62

kraatzii Heyden, 1882, Poecilus [Derus] 484



kraetschmeri Drovenik, 1978, Carabus 163

krajnensis Born, 1904, Carabus 165

krali Deuve, 1994, Carabus 176

krali Deuve, 1991, Cychrus 204

krali Farka, 1993, Leistus 80

krali Sciaky, 1997, Pterostichus 514

kralianus Deuve, 1996, Cychrus 204

krapog Morvan, 1999, Platyagonum 462

krasae Roubal, 1903, Carabus 130

krasnobaevi O. Berlov & Tilly, 1998, Pterostichus 506

krasnopolensis Kirschenhofer, 1987, Pterostichus 504

krasnovi Belousov & Kabak, 1992, Trechus 333

kratkyi Ganglbauer, 1890, Carabus 131

kratochvili Kult, 1949, Poecilus [Pterostichus] 483

kratteri Dejean, 1831, Nebria 92



kraussei Born, 1908, Carabus 152

kraussei Reitter, 1914, Typhloreicheia [Reicheia] 223



kraussi Winkler, 1923, Anophthalmus 291

kraussi Lapouge, 1898, Carabus 139

kraussi Ganglbauer, 1897 (as 1896), Trechus 339

kravetzi Komarov, 1993, Meganophthalmus 314



kreissli Schweiger, 1964, Carabus 148

krejcii Deuve & Quéinnec, 1985, Trechus 333



krekichi Schatzmayr, 1909, Amara 547

krestovyanus Vysoký, 1984, Pterostichus 496

kretaense Kirschenhofer, 1982, Agonum 450

kricheldorffi Puel, 1925, Badister 440

kricheldorffi Breuning, 1926, Carabus [Mesocarabus] 167

kricheldorffi Le Moult, 1913, Carabus 140

kricheldorffi Wagner, 1932, Chlaenius 352

kricheldorffi Fuente, 1921, Cymindis 413

kricheldorffi Wagner, 1913, Trechus 334



krivolutzkajae Lafer, 1989, Amara 557

krnensis Bernau, 1911, Carabus 184

kroli J. L. omnicki, 1903, Notiophilus 96

kronii Hoppe & Hornschuch, 1825, Carabus 171

kruberi Fischer von Waldheim, 1820, Carabus 191

kruberoides Breuning, 1966, Carabus 191

krueperi Apfelbeck, 1904, Amara 562

krueperi Reitter, 1896, Carabus [Chaetocarabus] 138

krueperi Schaum, 1862, Duvalius [Anophthalmus] 303



krueperi Reitter, 1884, Glycia [Neotarus] 409

krueperi L. Miller, 1884, Laemostenus 532

krueperi Apfelbeck, 1904, Lebia 427

krueperi Reitter, 1885, Omphreus 446

krueperi Apfelbeck, 1904, Ophonus [Harpalus] 388

krueperi Apfelbeck, 1904, Tachyura [Tachys] 280

krueperi V. Guéorguiev & B. Guéorguiev, 1999, Tapinopterus 520

krueperii Seidlitz, 1887, Gyrinus 27

krugeri Obydov, 1999, Carabus 172

krulikovskyi Lutshnik, 1933, Pterostichus [Platysma] 496

krynickii Hrdlika, nom. nov., Brachinus 19, 215



krynickii Fischer von Waldheim, 1828, Carabus 199

krynickii Sperk, 1835, Platynus [Anchomenus] 464

kryshanovskii Jeannel, 1962, Duvalius 304

kryshanowskii Hieke, 1976, Amara 560

kryshtali Bilyashivsky, 1993, Hydroporus 62

kryzhanovskianus Deuve, 1992, Carabus 125

kryzhanovskii Mikhailov, 1988, Bembidion 258

kryzhanovskii Belousov & Kabak, 1992, Brachinus 212

kryzhanovskii Mikhailov, 1970, Bronislavia 363

kryzhanovskii J. Schmidt, 1999, Calathus 525



kryzhanovskii Bogachev, 1965, Carabus 145

kryzhanovskii Zamotajlov & Sciaky, 1999, Chinapenetretus 281

kryzhanovskii Belousov, 1998, Cimmerites 297

kryzhanovskii Emetz, 1972, Cymindis 416

kryzhanovskii Zamotajlov, 1999, Deltomerodes 281

kryzhanovskii Zamotajlov, 1988, Deltomerus 282

kryzhanovskii Gryuntal, 1984, Dyschirius 230

kryzhanovskii Kataev, 1988, Harpalus 378

kryzhanovskii Shilenkov, 1982, Nebria 88

kryzhanovskii Wrase, 1999, Oedesis 366



kryzhanovskii Vereschagina, 1988, Pseudotaphoxenus [Taphoxenus] 541

kryzhanovskii Lindroth, 1966, Stereocerus [Pterostichus] 517

kryzhanovskii Lafer, 1989, Trechiama [Leptepaphiama] 321

kryzhanovskii Kataev & N. Ito, 1999, Trichotichnus 394

kryzhanovskiji Sciaky & Facchini, 1999, Loricera 98

kryzhanovskiji Casale, 1988, Pseudotaphoxenus 541

kuangcensis Mlyná, 1974, Harpalus 384

kuatensis Jedlika, 1956, Amara 567

kubanensis Lutshnik, 1921, Nebria 87

kubani Deuve, 1990, Carabus 142

kubani Deuve, 1992, Cychrus 204

kubani Farka, 1993, Leistus 80

kubaniana Ledoux & Roux, 1998, Nebria [add to incertae sedis] 95

kubanicus Belousov, 1998, Apocimmerites 295



kubanicus Reitter, 1896, Carabus 131

kubanicus Jedlika, 1957, Harpalus 372

kubanicus Kirschenhofer, 1987, Pterostichus [HN, repl. in vol. 2] 504



kubanicus Reitter, 1896, Pterostichus 496

kubaniensis Semenov, 1897, Carabus 200

kubotai Uéno, 1958, Stygiotrechus 320

kucerai Deuve, 1997, Carabus 128

kucerai Deuve, 1997, Cychrus 203

kucerai Farka, 1995, Leistus 80

kucerai Sciaky, 1997, Pterostichus 500

kuchtae Breit, 1909, Bembidion 267

kuchtai Breit, 1908, Graptodytes [Hydroporus] 59

kuciensis Nonveiller & Pavievi, 1987, Winklerites 240

kuenlunensis Bates, 1878, Amara [Amathitis] 562

kuennemanni Langhenhan, 1913, Carabus 195

kuesteri Jakobson, 1906, Bembidion 253

kuesteri Schaum, 1845, Bembidion 243



kuesteri Csiki, 1906, Cymindis 412

kuesteri Schaufuss, 1865, Laemostenus [Antisphodrus] 532

kugelanni Panzer, 1797, Poecilus [Carabus] 483

kugitangensis Deuve & Kaláb, 1993, Carabus 134


kuhitangi Mikhailov & Belousov, 1991, Bembidion 256

kuhitangi Kryzhanovskij, 1962, Chilotomus 364

kukunorensis Semenov, 1898, Carabus 198

kuldshensis Ballion, 1878, Carabus 191

kuliabense Netolitzky, 1931, Bembidion 257

kulingensis Kleinfeld & Schütze, 1997, Carabus 156

kuljabensis Jedlika, 1961, Pseudotaphoxenus [Taphoxenus] 541

kulleri Vondel, 1988, Haliplus 32

kulpensis Belousov & Kabak, 1998, Trechus 334

kulti Fassati, 1947, Amara 567

kulti Fassati, 1942, Bembidion 259

kulti Jedlika, 1952, Colpoides 457

kulti Jedlika, 1960, Desera 358



kulti Jedlika, 1963, Desera 358

kulti Jedlika, 1947, Dromius [Lebidromius] 421



kulti Roubal, 1948, Dyschirius 229

kulti Maan, 1939, Nebria 85

kulti Jedlika, 1940, Platynus [Colpodes] 465

kulti Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

kulti Maan, 1940, Tapinopterus 520

kultianus Farka & Fassati, 1999, Leistus 83

kultianus Jedlika, 1946, Pterostichus 491

kultukus Jedlika, 1958, Pterostichus 508

kuluensis Bates, 1891, Chlaenius 350

kuluensis Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

kulzeri Netolitzky, 1935, Bembidion 259



kulzeri Breuning, 1932, Carabus 134

kulzeri Schauberger, 1934, Carterus 364

kulzeri Straneo, 1942, Pterostichus 501

kumagaii Kimura & Komiya, 1974, Carabus [Leptocarabus] 159

kumanensis Reitter, 1884, Tapinopterus [Pterostichus] 518

kumaso Imura & Mizusawa, 1995, Carabus 177

kumatai Habu, 1973, Bembidion 20, 269

kumatai Habu, 1973, Leistus 80

kumatai Habu, 1979, Orthogonius 446

kumatai Habu, 1973, Tostkar [Agonum] 467

kumatai Habu, 1973, Xestopus [Calathus] 530

kumataianum Habu, 1973, Agonum 454

kumensis Fischhuber, 1975, Anophthalmus 292

kumgangi Shilenkov, 1983, Nebria 88

kunarense Kirschenhofer, 1989, Bembidion 269

kunarensis Kataev, 1993, Harpalus 378

kunashirensis Pütz & Wiesner, 1995, Cylindera 110

kungeicus Breuning, 1934, Carabus [Trachycarabus] 146

kungeicus Shilenkov & Kabak, 1994, Leistus 82



kuniakii Ishikawa, 1971, Carabus [Procrustes] 164

kunikanei Nakajima, 1993, Carabus [Acoptolabrus] 122

kunmingensis Kirschenhofer, 1996, Parena 434



kunstleri Peytoureau, 1894, Dytiscus 51

kuntzei Polentz, 1939, Dromius 420



kuntzeni Eidam, 1930, Carabus 195

kuntzeni Dupuis, 1914, Pheropsophus 217

kuntzeni J. Müller, 1932, Tapinopterus 520



kunzei Heer, 1837, Carabus 166

kunzei Gistel, 1838, Cylindera [Cicindela] 109

kuporosovi E. Berlov & O. Berlov, 1996, Pterostichus 505

kuqaensis Deuve, 1994, Carabus 160

kuraicus Shilenkov, 1995, Trechus 334

kuraminensis Kabak, 1997, Cymindis 416

kuraminensis Vereschagina & Kabak, 1997, Laemostenus 537

kurangense Morvan, 1973, Bembidion [Emphanes] 248

kurbatovi Kataev & N. Ito, 1999, Trichotichnus 394

kurdaiensis Vereschagina & Kabak, 1996, Pseudotaphoxenus 541

kurdaorum Lassalle, 1992, Carabus 193

kurdicus Heinz, 1975, Carabus 193

kurdicus Casale, 1988, Laemostenus 534



kurdistanicola Deuve, 1990, Carabus 198

kurdistanicum Netolitzky, 1920, Bembidion 256

kurdistanicus Eidam, 1931, Carabus [Cyclocarabus] 198

kurdistanicus P. Machard, 1986, Carabus 157

kurdistanicus Mateu, 1961, Microlestes 423

kurdistanicus Freude, 1989, Zabrus 571

kurdistanus Casale, 1988, Laemostenus 538

kurdistanus Schauberger, 1932, Pachycarus 367

kurentzovi Lafer, 1989, Nebria 89

kurentzovi Lafer, 1979, Pterostichus 495

kurentzovi Lafer, 1989, Trechus 334



kurilensis Kamiya, 1938, Agabus 41

kurilensis Lapouge, 1913, Carabus 134

kurilensis Lafer, 1989, Dromius 420

kurnakovi Budarin & Kryzhanovskij, 1995, Amara [Curtonotus] 560

kurnakovi Kryzhavoskij, 1983, Anchomenus [Agonum] 455



kurnakovi Schuler, 1962, Bembidion [Peryphus] 255

kurnakovi Kryzhanovskij, 1968, Carabus 186

kurnakovi Jeannel, 1960, Cimmerites 297

kurnakovi Zamotajlov, 1988, Deltomerus 283

kurnakovi Jeannel, 1960, Duvalius 307

kurnakovi Dolzhanski & Ljovuschkin, 1989, Inotrechus 312

kurnakovi G. E. Ball, 1966, Pterostichus 491

kurnakovi Kryzhanovskij, 1988, Pterostichus [HN] 505


kurnakovi Jeannel, 1960, Trechus 334

kurnakowi Hieke, 1994, Amara 560

kurokawense Habu & Baba, 1958, Bembidion 265

kurokawensis Habu & Baba, 1957, Pterostichus 510

kurosai Habu, 1967, Parena 434

kurosai Tanaka, 1958, Pterostichus 491

kurosai Uéno, 1959, Rakantrechus 319

kurosai Tanaka, 1962, Stenolophus 405

kurosai Uéno, 1956, Thalassoduvalius 321

kurosai Uéno, 1955, Trephionus 546

kurosai Habu, 1973, Trichotichnus 396

kurosawai Morita, 1986, Apatrobus 280

kurosawai Breuning, 1957, Carabus 149

kurosawai Kasahara, 1986, Chlaenius 348



kurosawai Satô, 1979, Lacconectus 48

kurosawai Nakane, 1960, Nebria 92

kurosawai Tanaka, 1958, Pterostichus 491

kurosawai Satô, 1982, Satonius [Delevea] 25

kurosawai Miyatake, 1986, Tenomerga 25

kurosawai Uéno, 1986, Trechiama 322

kurosonense Habu, 1957, Agonum 454

kurosonis Kasahara, 1994, Pterostichus 512

kurram Andrewes, 1935, Bembidion 258

kurucanicum Kirschenhofer, 1981, Agonum 451

kuruschensis Franz, 1991, Trechus 335

kuruschicum Netolitzky, 1930, Bembidion 255

kusakarii Uéno, 1986, Trechiama 322

kusamai Uéno, 1999, Ishikawatrechus 312

kusamai Habu, 1977, Stenolophus 405

kuschakewitschi Ballion, 1871, Calosoma [Callisthenes] 119



kusdasi Mandl, 1966, Carabus 185

kushironis Uéno, 1992, Trechus 325



kusnetzovi Shilenkov, 1996, Carabus 133

kusnetzowi Semenov, 1903, Carabus 153



kusteri Marseul, 1866, Laccophilus 76

kutaissianus Zaitzev, 1941, Meganophthalmus [Duvalius] 314



kutensis Poppius, 1905, Pterostichus 507

kutschumi Putchkov, 1993, Cephalota [Cicindela] 102

kutsherai Sciaky & Wrase, 1997, Myas [Trigonognatha] 476

kuzminae Shilenkov, 1990, Carabus 174

kuznetsovi Uéno & Lafer, 1992, Trechiama 322

kuznetzovi Lafer, 1989, Amara 567

kuznetzovi Lutshnik, 1928, Amara 559

kuznetzovi Lafer, 1989, Bradycellus 402

kuzonduresti Morvan, 1995, Xestagonum [Colpodes] 468



kuzugamii Shibata, 1987, Dolichoctis 436

kvirensis Belousov, 1992, Carabus 181

kvirensis Belousov, 1991, Pterostichus 504

kwanmobongensis Kwon & Lee, 1984, Carabus [Leptocarabus] 134

kweitshauensis Mandl, 1975, Carabus 128

kwonileeique Deuve, 1992, Carabus 159

kykladensis Breuning, 1961, Carabus 182

kyllenicus Scheibel, 1937, Duvalius 304

kyokoae Uéno & Baba, 1974, Kurasawatrechus 313

kyparissis Jedlika, 1963, Tapinopterus [Pterostichus] 520

kyros Netolitzky, 1931, Bembidion 257

kyushuense Habu, 1954, Agonum 452

kyushuense Habu, 1957, Bembidion 267

kyushuensis Nakane, 1961, Carabus 159

kyushuensis Nakane, 1990, Leiodytes [Clypeodytes] 56

kyushuensis Habu, 1958, Nippononebria 95

kyushuensis Habu, 1967, Pentagonica 448

kyushuensis Habu, 1955, Pterostichus 512

kyushuensis Habu, 1978, Trephionus 547

labacensis J. Müller, 1931, Anophthalmus 293

labarthei A. Gaudin, 1947, Aphoenops 295

labensis Belousov, 1998, Alanorites 290



labensis Gottwald, 1985, Carabus 132

labiatus Brahm, 1790 (as 1791), Agabus [Dytiscus] 37



labiatus Motschulsky, 1849, Daptus 369

labionigra Mandl, 1965, Calochroa [Cicindela] 99



labittei Clément, 1904, Carabus 194

labradorensis Casey, 1918, Amara [Curtonotus] 561

labradorensis Fall, 1923, Hydroporus 62

labradorinus Casey, 1918, Patrobus 285

labrang Farka, 1999, Leistus 80

labrangicus Deuve, 1992, Carabus 189

labroturrita Cassola & Wranik, 1998, Socotrana 115

labruleriei Géhin, 1884, Carabus [Chaetomelas] 188

labruleriei Jeannel, 1921, Trechus 341

labzuki Lafer, 1979, Pterostichus 495

laccophiloides A. Zimmermann, 1928, Lacconectus 48

laccophilus Reitter, 1896, Carabus 160

lacerans Bates, 1889, Pristosia [Pristodactyla] 531



laceratus Csiki, 1927, Carabus 199

lacertosus Sturm, 1815, Badister 440



lacertosus Dejean, 1828, Percus [Feronia] 478

laconicus Cerruti, 1973, Speluncarius 516



lacordairei Putzeys, 1846, Pterostichus [Steropus] 514

lacouri Antoine, 1941, Orthomus [Platysma] 477



lacouri Antoine, 1957, Zabrus 572

lacrimans Netolitzky, 1935, Bembidion 259



lacrymosus Lapouge, 1899, Carabus 152

lactarius Uéno, 1959, Rakantrechus 319

lacteola Pallas, 1776, Cicindela 106

lacunosus Andrewes, 1920, Chlaenius 352

lacunosus Chaudoir, 1844, Pterostichus [Feronia] 500

lacustrina Casey, 1918, Amara 549

lacustris L. Redtenbacher, 1858, Dicheirotrichus [Bradycellus] 402



lacustris Panzer, 1796, Ilybius [Dytiscus] 42

lacustris Gistel, 1857, Lionychus 432

lacustris Motschulsky, 1844, Patrobus 285

ladakense Bates, 1878, Agonum [Anchomenus] 450

ladakense Andrewes, 1924, Bembidion 261

ladakensis Andrewes, 1929, Dyschirius 226

ladakhensis Kirschenhofer, 1992, Trichotichnus 396

ladas Andrewes, 1924, Bembidion 243

ladikensis Deuve & Charet, 1992, Carabus 158



ladinicus J. Müller, 1951, Laemostenus 535

ladygini Semenov, 1903, Carabus 190



laemostenomimus Lutshnik, 1915, Poecilus [Platysma] 482

laesipennis Bates, 1873, Parena [Crossoglossa] 434



laeta Motschulsky, 1844, Cylindera [Cicindela] 109

laetabilis Walker, 1858, Canthydrus [Hydroporus] 34



laetecupreoviridis Chaudoir, 1865, Calomera [Cicindela] 99

laetescripta Motschulsky, 1860, Chaetodera [Cicindela] 103

laeticolor Bates, 1873, Bradycellus 401

laeticulus Sharp, 1882, Graptodytes [Hydroporus] 60

laetifica Bates, 1873, Tachyura [Tachys] 279



laetifrons J. Müller, 1941, Crasodactylus 362

laetiusculus Chaudoir, 1856, Chlaenius 350

laetula Tschitschérine, 1903, Cylindera [Cicindela] 111

laetula Reitter, 1893, Cymindis 415

laetula Bates, 1889, Dioryche [Platymetopus] 369


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