Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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prenta Jedlika, 1937, Pristosia [Calathus] 531

prentus Jedlika, 1937, Calathus 528


prepyrenaeus Coiffait, 1974, Trechus 332

prescotti Mannerheim, 1829, Ilybius [Colymbetes] 42

preslicensis Apfelbeck, 1904, Molops 474

preslii Dejean, 1830, Carabus 179

pressicollis Cavazzuti, 1999, Carabus 189



pretiosissimus G. Hauser, 1921, Carabus [Coptolabrus] 143

pretiosum Friederichs, 1903, Agonum [Platynus] 450

pretiosus Rosenhauer, 1856, Chlaenius 352

pretiosus Faldermann, 1837, Laemostenus [Pristonychus] 538

pretneri Scheibel, 1935, Adriaphaenops 289

pretneri J. Müller, 1913, Anophthalmus [Trechus] 292



pretneri Krätschmer & Drovenik, 1977, Carabus 163

pretneri V. B. Guéorguiev, 1971, Duvalius 308

pretneri Scheibel, 1936, Neotrechus 316

pretneri Meggiolaro, 1957, Orotrechus 317

prevosti Gory, 1833, Carabus 157

prevostii Dejean, 1828, Pterostichus [Feronia] 488



preyssleri Duftschmid, 1812, Carabus 173

priamus Breuning, 1948, Carabus 125

priapus K. Daniel, 1902, Trechus 338



prichodkoi Semenov, 1897, Carabus 137

priesneri Schauberger, 1921, Speluncarius 517

priesneri Schatzmayr & Koch, 1934, Tachyura [Tachys] 21, 279

primarius Lapouge, 1902, Carabus 163

primigenioides Jeanne, 1974, Trechus 331

primigenius J. Müller, 1919, Orotrechus [Trechus] 317



primigenius Jeannel, 1920, Trechus 23, 331

primitiva Jedlika, 1957, Amara 559

princeps Nietner, 1857, Chlaenius 349

princeps Chaudoir, 1852, Lebia 430

princeps Bates, 1883, Myas [Trigonognatha] 476

princeps Sharp, 1882, Nebrioporus [Deronectes] 66

princeps Régimbart, 1888, Platambus [Platynectes] 44

principalis Bates, 1889, Carabus 143



principatus Schaufuss, 1872, Carabus 173

prinsepii Saunders, 1834, Cylindera [Cicindela] 111

priscus J. Müller, 1922, Dyschirius 230



pristinus Friederichs, 1903, Notiophilus 96

problematicior Deuve, 1994, Carabus 160

problematicus Herbst, 1786, Carabus 168

problematicus Bokor, 1913, Duvalius [Anophthalmus] 300

proboscideus Olivier, 1795, Cychrus [Carabus] 202

probsti Deuve, 1991, Carabus 162

probsti Mandl, 1976, Cephalota [Cicindela] 103

probsti Kirschenhofer, 1996, Dischissus 447



probsti Kirschenhofer, 1994, Peliocypas [Risophilus] 434

probsti Kirschenhofer, 1992, Trichotichnus 395

probus Park & Kwon, 1996, Pterostichus 495

procax A. Morawitz, 1862, Pterostichus 495



procedens Csiki, 1927, Carabus 168

procephalus Bates, 1873, Pterostichus 487

proceroides Jeannel, 1926, Duvalius 307

procerulus Chaudoir, 1862, Carabus 159

procerulus Heyden, 1880, Pterostichus [Haptoderus] 492

procerus Régimbart, 1899, Agabus [Platynectes] 36

procerus Bielz, 1887, Carabus 151

procerus Reitter, 1885, Carabus 163

procerus Uéno, 1982, Chaetotrechiama 297



procerus Sharp, 1882, Colymbetes 45

procerus Cavazzuti, 1998, Cychrus 205

procerus Putzeys, 1847, Duvalius [Trechus] 307

procerus Joseph, 1870, Laemostenus [Sphodrus] 535

procerus Schaum, 1858, Loxoncus [Stenolophus] 404

procerus Dejean, 1825, Scarites 234

procerus Aubé, 1838, Stictotarsus [Hydroporus] 69

processifer Belousov & Kabak, 1992, Trechus 338

prochazkai Obenberger, 1913, Typhlotrechus [Anophthalmus] 345



prochazkorum Jedlika, 1967, Amara 557

prochazkorum Jedlika, 1967, Pterostichus 506

procrustoides Géhin, 1885, Carabus [Pachystus] 158

proderus Fairmaire, 1880, Paradromius [Dromius] 424

prodigus Erichson, 1834, Carabus 126



productum Schilsky, 1889, Bembidion 248

productus Hampe, 1852, Carabus 158

productus Fairmaire, 1880, Hydroporus 64

productus Régimbart, 1884, Orectochilus 29

proelongus Reiche & Saulcy, 1855, Orthomus [Feronia] 477

professoris Uéno, 1999, Ishikawatrechus 312



profundecrenatus Wollaston, 1867, Poecilus [Pterostichus] 480

profundestriatus Schatzmayr, 1937, Paradromius [Dromius] 424



profundestriatus Heer, 1837, Trechus 331

profundissimus Mihók, 1914, Duvalius [Trechus] 300

profundus Mlejnek & J. Moravec, 1996, Aphaenopidius 293

prognathus Bates, 1883, Stomis 517

progrediens Semenov, 1897, Carabus 192

progrediens Schauberger, 1922, Harpalus 380



progressus Lapouge, 1925, Carabus 168

progressus Semenov, 1888, Carabus 178

prokletiensis Horvatovich, 1974, Nebria 85

prokletijensis Maan, 1938, Calathus 527


prokopenkoi Mikhailov, 1995, Bembidion [add] 250

prokschi Mader, 1917, Brachinus 214

proksi Jedlika, 1963, Coptodera 436

prolapsus Cavazzuti, 1999, Cychrus 205

prolixa Kiesenwetter, 1857, Clivina 219

prolixa Bates, 1873, Lachnocrepis [Oodes] 445

prolixus Bates, 1883, Dromius 421

prolixus Fairmaire, 1875, Laemostenus [Sphodrus] 535



prolixus Putzeys, 1876, Poecilus 483

prolixus Bates, 1892, Tachys 276

prolongatus Bates, 1883, Leistus 83

prolongatus A. Morawitz, 1862, Pterostichus 493

promachus Bates, 1891, Carabus 163

prometheus Reitter, 1887, Carabus 132



prominulum Puel, 1938, Agonum 449

promissus Reiche, 1872, Graniger [Harpalus] 366

promissus Heyden, 1875, Molops 474



promota Motschulsky, 1866, Nebria 94

pronepta Ishikawa, 1992, Carabus [Leptocarabus] 159

pronotalis W. Horn, 1922, Cylindera 111

pronotalis W. Horn, 1901, Neocollyris [Collyris] 116

propagator Reitter, 1901, Eocarterus [Odontocarus] 366



properans C. Zimmermann, 1832, Amara 556

properans Stephens, 1828, Bembidion [Tachypus] 251

properans Chaudoir, 1868, Pterostichus [Feronia] 492

propevau Fassati, 1944, Bembidion 263

propheta Rapuzzi, 1995, Carabus 157



prophettei Reiche, 1862, Apristus 431

prophylax Heller, 1903, Pheropsophus 218

propinqua Ménétriés, 1832, Amara 561

propinqua Chaudoir, 1835, Cephalota [Cicindela] 102

propinquus Ballion, 1871, Anisodactylus 361

propinquus Csiki, 1927, Carabus 185



propinquus Joseph, 1870, Laemostenus [Sphodrus] 535

propinquus A. Morawitz, 1862, Stenolophus 406



propinquus Chaudoir, 1844, Zabrus 572

proserpina Jeannel, 1909, Aphoenops 294



proserpina Mandl, 1955, Carabus 148

proserpina Normand, 1939, Reicheia 222



proserpinae Ragusa, 1886, Acupalpus 399

prosper Lapouge, 1918, Carabus 192

prostenus Bates, 1873, Chlaenius 350

prosternalis Ganglbauer, 1891, Oreonebria [Nebria] 96

prostratum Motschulsky, 1844, Bembidion [Notaphus] 248

protalmum Netolitzky, 1934, Bembidion 247


protectus Winkler, 1926, Duvalius 300

protenes Bates, 1889, Carabus 126



protensa Motschulsky, 1844, Nebria 94

protensoides Novotný & Voíšek, 1988, Carabus 132

protensus Schaum, 1864, Carabus 132

protensus Putzeys, 1866, Dyschirius 226

protensus A. Morawitz, 1863, Platynus [Dyscolus] 463

protensus Rosenhauer, 1856, Platytarus [Cymindis] 410

protensus Schaum, 1857, Tapinopterus [Feronia] 520

proteron Netolitzky, 1920, Bembidion 271

proteus Paykull, 1790, Harpalus [Carabus] 372

proteus Régimbart, 1877, Laccophilus 77

provincialis Born, 1903, Carabus 165

provincialis Puel, 1925, Dyschirius 225

provincialis Carret, 1903, Pogonus 287

provincialis Jeannel, 1942, Trichotichnus 396



provoanum Casey, 1918, Bembidion 254

provostii Fairmaire, 1888, Carabus 135

proxima J. Frivaldszky, 1877, Amara 551

proxima Putzeys, 1866, Amara 551

proxima A. Morawitz, 1862, Pristosia [Calathus] 531

proxima Chaudoir, 1860, Prothyma [Odontochila] 115



proximatoides Depoli, 1925, Carabus 198

proximatus Reitter, 1896, Carabus 198

proximior Deuve, 1997, Carabus 191



proximum Aubry, 1980, Bembidion [Ocydromus] 260

proximus Sharp, 1882, Canthydrus 34



proximus Reitter, 1885, Carabus [Procrustes] 187

proximus Martínez y Saez, 1873, Chlaenius 353



proximus LeConte, 1848, Harpalus 379

proximus Stephens, 1828, Hydroporus 64

proximus J. Frivaldszky, 1879, Platynus [Anchomenus] 463

proximus Dejean, 1829, Stenolophus 406

pruinosus Jeannel, 1937, Duvalius 308



prunicolor Deuve, 1996, Carabus 189

prunieri Pham, 1978, Carabus 168

prunierorum Lassalle, 1998, Omphreus 446



prunneri Mallász, 1900, Carabus 151

pruthii Vazirani, 1966, Haliplus 32



pryeri Bates, 1883, Metacolpodes [Colpodes] 460

prymnaeus Fischer von Waldheim, 1828, Cychrus 203

przewalskensis Deuve, 1992, Carabus 123

przewalskii Semenov, 1889, Broscus 236

przewalskii A. Morawitz, 1886, Carabus 135

przewalskii Dokhtouroff, 1887, Cicindela 104



przewalskii Semenov, 1889, Diplous [Platidius] 283

przewalskii Semenov, 1889, Discoptera 356

przewalskii Semenov, 1889, Nebria 91

przewalskii Semenov, 1889, Pristosia 531

przewalskii Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 540

przewalskii Semenov, 1889, Zabrus 571



przewalskyi Tschitschérine, 1894, Harpalus 383

przewalskyi Tschitschérine, 1888, Pterostichus 514

przewalskyi Belousov & Kabak, 1993, Trechus 338

psakhoensis Belousov, 1998, Porocimmerites 318

psammodes Andrewes, 1932, Cymindis 414

psammodes P. Rossi, 1792, Nebria [Carabus] 91

psammodroma Chevrolat, 1845, Abroscelis [Cicindela] 99

psammophila Solsky, 1874, Nebria 91

pschadensis Zamotajlov, 1990, Carabus 132

pseudagnatus Novotný & Voíšek, 1988, Carabus 200

pseudamphinomus Habu, 1974, Platyagonum [Platynus] 462

pseudapostolus Schaufuss, 1865, Laemostenus [Antisphodrus] 532

pseudescherichi Breuning, 1962, Carabus 158



pseudextensus A. Fleischer, 1899, Dyschirius 224

pseudinsulicola Ishikawa, 1966, Carabus [Apotomopterus] 177



pseudoacuticollis atloš & Moravec, 1999, Carabus 188

pseudoaeneolus Schauberger, 1930, Harpalus 372

pseudoaeneus Dejean, 1829, Anisodactylus 361

pseudoalmonius Deuve, 1993, Trechus 338

pseudoalpestris Mlyná, 1977, Molops 474



pseudoalpicola Maan, 1941, Leistus 18, 82

pseudoalyshensis Deuve & Quéinnec, 1992, Trechus 338



pseudoaxillaris Bedel, 1906, Cymindis 414

pseudoazureus Reitter, 1900, Ophonus 389

pseudobarthei Schaefer, 1973, Carabus [Autocarabus] 139

pseudobliteratus Korell & Kleinfeld, 1982, Carabus 200



pseudoboeticus Lassalle, 1986, Carabus 161

pseudobonellii Lapouge, 1903, Carabus 184

pseudobruggei Deuve, 1991, Cychrus 203

pseudobrunnea Lindroth, 1968, Amara 557

pseudobrunneus Casale, 1981, Laemostenus 538

pseudocallosum Paul Meyer, 1949, Bembidion 253

pseudocameroni Deuve, 1996, Trechus 338



pseudocantalicus Méquignon, 1914, Pterostichus 512

pseudocarabus Semenov & Redikorzev, 1928, Calosoma 119

pseudocareniger Deuve, 1991, Carabus 133

pseudocarpathicum Netolitzky, 1943, Bembidion 259



pseudocaucasica Mandl, 1955, Calomera [Cicindela] 100

pseudochlorothyreus G. Hauser, 1921, Carabus [Coptolabrus] 143

pseudocholaensis Deuve, 1998, Trechus 338

pseudocinctulus Mandl, 1961, Ropaloteres [Cicindela] 114

pseudocinctus Mandl, 1960, Ropaloteres [Cicindela] 114

pseudoclypealis Scholz, 1933, Agabus 36

pseudocoelestis Kleinfeld, 1999, Carabus 143

pseudocoiffaitianus Deuve, 1990, Carabus 149

pseudocollaris Casale, 1988, Pseudotaphoxenus 542



pseudocolossus Cavazzuti, 1993, Carabus 158

pseudocommunis Burakowski, 1957, Amara 551

pseudoconfertus Wallis, 1926, Ilybius [Agabus] 42

pseudoconfluens Sawada & Wiesner, 1999, Therates 116

pseudoconsumatum Kirschenhofer, 1984, Bembidion 268

pseudocorporosus Schauberger, 1930, Harpalus [Haploharpalus] 383

pseudocratophyrtus Obydov, 1998, Carabus 145

pseudocrenifer Sciaky, 1995, Leistus 81

pseudocruciatus Reitter, 1909, Brachinus 215

pseudocrypticola Coiffait, 1947, Aphoenops 294

pseudocyaneum Belousov & Sokolov, 1994, Bembidion 246

pseudodepressus P. Machard, 1988, Carabus 123

pseudodeserticola W. Horn, 1891, Cylindera [Cicindela] 111

pseudodiamesus Ivanovs, 1994, Carabus 134



pseudodimidiatus Schauberger, 1928, Harpalus 374

pseudodolens Morvan, 1978, Pterostichus 510

pseudoducalis Heinz & Korge, 1967, Carabus 158

pseudoelongatus P. Machard, 1994, Carabus 193

pseudoerosus Mandl, 1955, Carabus 192



pseudoerosus Mandl, 1967, Carabus 201

pseudofaustulus Le Moult, 1912, Carabus 140

pseudofemoralis F. Battoni, 1986, Calathus 527



pseudofischeri Mandl, 1972, Calomera [Cicindela] 100

pseudoflavus Rocchi, 1986, Canthydrus 34

pseudofossiger Stark, 1894, Carabus 200

pseudofulgens Born, 1905, Carabus 165

pseudofulva Ali, 1967, Amara 566

pseudogandharae Deuve, 1995, Carabus 155

pseudogebleri Deuve, 1994, Carabus 174

pseudogotschi Breuning, 1926, Carabus 131



pseudogracilior Panin, 1943, Carabus 197

pseudograniger Reitter, 1896, Carabus 196

pseudogranulatus Nüssler, 1965, Carabus 136

pseudogratus Deuve, 1994, Carabus 135

pseudoguzhangensis Cavazzuti & Ratti, 1999, Carabus 125

pseudoharmandi Mandl, 1965, Carabus 162

pseudoharmandi Morvan, 1981, Pterostichus 510

pseudoheros Kenyery, 1983, Carabus 184


pseudohuangi Deuve, 1995, Carabus 147

pseudoignigena Kleinfeld, 1996, Carabus 143

pseudoindigestus Deuve & Mourzine, 1998, Carabus 190

pseudojucundus Mandl, 1986, Pristomachaerus 356

pseudokaratavensis Deuve & Kaláb, 1993, Carabus 147

pseudokarelini Mandl, 1954, Calosoma 119

pseudokoiwayai Deuve & Mourzine, 1998, Cychrus 203

pseudokolatensis Heinz & Korge, 1967, Carabus 193

pseudokoreanus Breuning, 1932, Carabus 134



pseudokrueperi Schauberger, 1933, Ophonus 389

pseudolampros Schweiger, 1964, Bembidion [Metallina] 251

pseudolamprostus Kaláb, 1996, Carabus 147

pseudolaterale Netolitzky, 1943, Bembidion 252

pseudolatipennis Deuve, 1991, Carabus 128



pseudoleachi Obydov, 1999, Carabus 164

pseudolitorea Ali, 1978, Cephalota [Cicindela] 102

pseudolongiceps Mandl, 1955, Carabus 137

pseudolucillum Netolitzky, 1938, Bembidion 271

pseudomacularis Schatzmayr, 1942, Cymindis 412

pseudomandli Probst & Wiesner, 1996, Therates 116



pseudomaritima Lengerken, 1912, Cicindela 106

pseudomarkamensis Deuve, 1992, Carabus 176

pseudomaroccana Roeschke, 1891, Cicindela 106

pseudomidas Mandl, 1967, Carabus 148

pseudominshanensis Deuve, 1992, Carabus 154

pseudomoestum Schuler, 1963, Agonum 451



pseudomoldaviensis Hormuzaki, 1907, Carabus [Morphocarabus] 170

pseudomontanellus Rizun, 1994, Trechus 338

pseudomonticola Lapouge, 1908, Carabus 130

pseudomorphus Semenov, 1889, Synuchus [Calathus] 546



pseudomuelleri Codina, 1921, Carabus 165

pseudonarentinus Mlyná, 1977, Molops 473



pseudoneesi Mandl, 1984, Carabus 166

pseudoneglectus Franciscolo, 1972, Ilybius [Agabus] 43



pseudonitens Muriaux, 1973, Trichotichnus 396

pseudonitidulum Reitter, 1908, Bembidion 259

pseudonivicola Kirschenhofer, 1983, Pterostichus 501

pseudonoricus Kenyery, 1983, Carabus 135

pseudoodes A. Fleischer, 1909, Harpalus 375

pseudoosmanilis Mlyná, 1977, Molops 474



pseudooxuroides G. Hauser, 1921, Carabus [Damaster] 148

pseudopachinus Nakane, 1963, Pterostichus 498



pseudopalustris Schulz, 1908, Cicindela 104

pseudopaphius Reitter, 1902, Ocys 273

pseudopappianus Imura & Yamaya, 1994, Carabus 133


pseudoparallelus Schauberger, 1926, Ophonus [Harpalus] 390

pseudoparis Deuve, 1992, Carabus 129

pseudoparoecus Csiki, 1905, Duvalius [Anophthalmus] 300

pseudopedius Reitter, 1887, Pterostichus 494



pseudopeltatus De Wailly, 1949, Badister 440

pseudopercus Reitter, 1889, Pterostichus [Aphaonus] 486

pseudophonoides Schauberger, 1930, Harpalus 385

pseudophrygius V. B. Guéorguiev, 1965, Graptodytes 60

pseudopiceus K. Daniel & J. Daniel, 1898, Trechus 338



pseudopictus Zaitzev, 1928, Graptodytes 59

pseudopilifer Straneo, 1957, Pterostichus [Trichomolops] 494

pseudopirithous Mandl, 1955, Carabus 181

pseudoplaga Netolitzky, 1943, Bembidion 249

pseudoplanipennis Mlyná, 1977, Molops 473

pseudoplatessa Gottwald, 1982, Carabus 200

pseudoplatyderus Sciaky, 1994, Pterostichus 500

pseudoplatynus Reitter, 1887, Deltomerus 283

pseudoplatysma Straneo, 1982, Pterostichus 510

pseudopolita Ledoux & Roux, 1996, Nebria 95



pseudoponticus Lassalle & P. Machard, 1991, Carabus 201

pseudopraestans Kurnakov, 1961, Lindrothius 529

pseudoprasinus Lapouge, 1912, Carabus 157

pseudoproperans Netolitzky, 1920, Bembidion 261

pseudoprosodes Semenov & Znojko, 1932, Carabus 147

pseudopshuensis Zamotajlov, 1991, Carabus 132

pseudopubescens A. Zimmermann, 1919, Hydroporus 64

pseudopurpurascens Kirschenhofer, 1987, Poecilus [Pterostichus] 485

pseudopusio Deuve, 1996, Carabus 190



pseudoquadricollis Schauberger, 1932, Ophonus [Harpalus] 391

pseudoremondianus Deuve, 1997, Carabus 189



pseudorichteri M. Machard & P. Machard, 1993, Carabus 161

pseudoriparia Mandl, 1935, Cicindela 107

pseudoripicola Iablokoff‑Khnzorian, 1963, Bembidion 250

pseudorobustus Heinz & Blumenthal, 1977, Carabus 157



pseudorugifer Sokolá, 1906, Carabus 187

pseudorugosus P. Machard, 1994, Carabus 131

pseudosackeni Deuve, 1989, Carabus 189



pseudosanchari Deuve & Mourzine, 1997, Carabus 176

pseudosaphyrinus Schweiger, 1965, Carabus 181



pseudoschweigerinae Schweiger, 1964, Carabus 193

pseudoscythicus Csiki, 1906, Carabus 197

pseudoseishini Hieke, 1990, Amara 559

pseudosemelai Cavazzuti, 1998, Carabus 126

pseudoserripes Reitter, 1900, Harpalus 380

pseudosiciliensis Dupuis, 1910, Cylindera [Cicindela] 111


pseudosiebkei Kirschenhofer, 1984, Bembidion 262

pseudosimplicidens Lafer, 1980, Amara 554

pseudosininensis Korell & Kleinfeld, 1988, Carabus 191


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