Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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saphyrinus Cristoforis & Jan, 1837, Carabus 181

saphyripennis Chaudoir, 1878, Platynus [Colpodes] 465



sapphirea Semenov, 1904, Cicindela 104

sapporense Jedlika, 1951, Bembidion 244

sapporensis Uchida & Tamanuki, 1927, Carabus [Acoptolabrus] 122

sapporoensis Jedlika, 1951, Lebia 430

saragaschensis Obydov, 1999, Carabus 174



sarajevoensis Apfelbeck, 1890, Carabus 163

sarajevoensis J. Müller, 1930, Molops 472

sarcoensis Knirsch, 1925, Duvalius [Trechus] 307

sardea Dejean, 1831, Lophyra [Cicindela] 113



sardeus Dejean, 1828, Laemostenus [Pristonychus] 539

sardeus Kuster, 1846, Laemostenus [Pristonychus] 537

sardingi A. Gaudin, 1947, Pterostichus [Lianoe] 498

sardiniensis A. Strand, 1917, Amara 551

sardoa Baudi di Selve, 1891, Typhloreicheia [Reicheia] 223

sardonius Jeannel, 1937, Anillus 238

sardoum Reitter, 1907, Omophron 208



sardous A. Fiori, 1903, Acupalpus 398

sardous Baudi di Selve, 1864, Amblystomus 360

sardous Dodero, 1917, Duvalius [Trechus] 305

sardous Jeannel, 1937, Hypotyphlus [Microtyphlus] 239

sardous Lucas, 1846, Laemostenus [Pristonychus] 537

sardous Baudi di Selve, 1883, Leistus 18, 84

sardous Holdhaus, 1912, Microlestes 422


sardous Küster, 1847, Olisthopus 461

sardous Schatzmayr, 1929, Poecilus [Pterostichus] 479

sardous Patrizi, 1955, Speomolops 517

sardus Gemminger & Harold, 1868, Nebrioporus [Hydroporus] 67

sareptanus K. Daniel, 1904, Harpalus [Ophonus] 374

sareptanus Jedlika, 1957, Poecilus [Pterostichus] 479

saridaghensis Shilenkov, 1983, Nebria 87

sarigoelensis Deuve & Charet, 1992, Carabus 194

sarikamisensis Cavazzuti, 1986, Carabus 194

sarkanduensis Deuve, 1981, Carabus 155

sarmaticus Bernau, 1913, Carabus 195

sarmaticus Motschulsky, 1850, Harpalus 375

sarmatus Schauberger, 1933, Ophonus [Harpalus] 392

sarrensis Laneyrie, 1948, Pterostichus [Troglorites] 514

sarsi Munster, 1927, Amara 550

saryarkense Kabak, 1992, Calosoma [Callisthenes] 118



sarydiazensis Deuve, 1992, Carabus 160

sarymsaktensis Obydov, 1999, Carabus 164

sasajii Uéno, 1980, Trechiama 323

sataense Habu, 1954, Agonum 449

sataensis Habu & Nakane, 1955, Trichotichnus 395

satanas Andrewes, 1924, Bembidion 246

satanas Semenov, 1897, Carabus 201

satanas Reitter, 1900, Harpalus 376

satie Vazirani, 1980, Nebrioporus [Potamonectes] 66

satoi Morita, 1993, Bembidion 252

satoi Wewalka, 1975, Hydaticus 54

satoi Löbl, 1994, Hydroscapha 25

satoi Wewalka, 1997, Microdytes 75

satoi Brancucci, 1982, Platambus 44

satoi Ishida, 1961, Pterostichus 507

satoi Habu, 1973, Stenolophus 405

satoi N. Ito, 1999, Trichotichnus 394

satoshii Ledoux, Roux & Sawada, 1991, Nebria 88

satoui Habu, 1976, Apatrobus [Patrobus] 280

satoui Uéno, 1976, Stygiotrechus 320

satoui Uéno, 1975, Trechiama 323

satsumanus Habu, 1958, Pterostichus 513

satsunanus Habu, 1974, Platynus 463

satunini Zaitzev, 1913, Ilybius [Agabus] 43

satunini Tschitschérine, 1903, Pterostichus [Platysma] 505



satuninianus Lutshnik, 1916, Pterostichus [Platysma] 487

saturatus Csiki, 1927, Carabus 143

satyrus Kurnakov, 1962, Carabus 132

satyrus Kurnakov, 1962, Pterostichus 505



satyrus Sturm, 1818, Trichotichnus [Harpalus] 396

saucius Desbrochers des Loges, 1871, Bidessus [Hydroporus] 55

saudiarabicus Mandl, 1980, Chlaenius 350

saudiarabicus Balkenohl, 1994, Dyschirius 227

saueri Deuve, 1992, Carabus 147

saueri Farka & P. Moravec, 1993, Duvalius 305

saueri Sciaky, 1994, Leistus 81

saueri Sciaky, 1995, Myas [Trigonognatha] 476

saueri Sciaky, 1994, Pterostichus 500

saueri Zamotajlov & Sciaky, 1996, Robustopenetretus [Parapenetretus] 286

saulcyanus Csiki, 1928, Trechus 340

saulcyi Piochard de la Brûlerie, 1875, Carabus 158

saulcyi Argod‑Vallon, 1913, Geotrechus [Trechus] 310

saulcyi Chevrolat, 1854, Macrocheilus 407

saulcyi Dieck, 1869, Scotodipnus 240

saulcyi Jeannel, 1921, Trechus 340

saundersi Andrewes, 1925, Elaphropus [Tachys] 273

saundersii Westwood, 1841, Paussus 210

saurica Shilenkov, 1976, Nebria 94

sauricus Belousov & Kabak, 1992, Trechus 340

sauteri Born, 1909, Calosoma 119

sauteri Roeschke, 1912, Carabus 127

sauteri Dupuis, 1914, Catascopus 435

sauteri Jedlika, 1954, Cillenus [Armatocillenus] 272

sauteri W. Horn, 1912, Cylindera [Cicindela] 110

sauteri Uyttenboogaart, 1915, Dineutus 26

sauteri A. Zimmermann, 1924, Haliplus 31

sauteri Dupuis, 1912, Hexagonia 407

sauteri Kuntzen 1911, Lesticus 472

sauteri Liebke, 1933, Mimocolliuris [Colliuris] 444

sauteri Wasmann, 1912, Paussus 210

sauteri Jedlika, 1956, Pogonus 287

sauteri W. Horn, 1912, Protocollyris [Collyris] 117

sauteri W. Horn, 1912, Therates 115

savinensis A. Zimmermann, 1933, Scarodytes [Deronectes] 69

savinicus Hammer, 1906, Carabus [Megodontus] 166

savinjscensis Daffner, 1996, Anophthalmus 22, 292

sawadai Hieke, 1994, Amara 564

sawadai Nakane, 1979, Nippononebria 95

sawadai Kamiya, 1932, Platambus [Agabus] 44

saxatile Gyllenhal, 1827, Bembidion 255

saxeseni Endrulat & Tessien, 1854, Cicindela 104

saxicola C. Zimmermann, 1832, Amara 565

saxicola Dejean, 1829, Harpalus 381

saxicola Tschitschérine, 1899, Pterostichus [Platysma] 497



saxicola Bonelli, 1813, Scarites 233

saxicola Putzeys, 1870, Trechus 340



sazonovi Cavazzuti, 1999, Carabus 188

sbordonii Vigna Taglianti, Genest & Sciaky, 1980, Duvalius 305

sbordonii Vigna Taglianti, 1967, Trechus 340

scaber Schaufuss, 1882, Carabus 166

scabra Fleutiaux, 1920, Tricondyla 118



scabridus Eidam, 1941, Carabus 197

scabripectus Huang, Hu & Sun, 1994, Harpalus 385



scabripenne Chaudoir, 1869, Calosoma 121

scabripennis Chaudoir, 1850, Carabus 198

scabriusculus Olivier, 1795, Carabus 199

scabrosus Olivier, 1790, Carabus 185



scalaris Audinet‑Serville, 1821, Cylindera [Cicindela] 110

scalesianus Stephens, 1828, Hydroporus 64



scalesii Stephens, 1828, Pedius [Argutor] 478

scalptus Sciaky & Wrase, 1997, Pterostichus 514

scandicum Lindroth, 1943, Bembidion 268

scandinavicus Born, 1926, Carabus [Mesocarabus] 168

scaphelytra Kavanaugh & Shilenkov, 1996, Nebria 89



scaphiformis Sharp, 1871, Hydroporus 63

scaphula Sturm, 1835, Hydroporus 64

scapulare Dejean, 1831, Bembidion 250



scapularis Stephens, 1828, Badister [Trimorphus] 441

scapularis Schaum, 1857, Cymindis 414

scapularis Geoffroy, 1785, Lebia [Buprestis] 429

scapularis Dejean, 1830, Mesolestes [Dromius] 421

scapulatus Belousov & Kabak, 1993, Trechus 340

scaritides Sturm, 1818, Pangus [Harpalus] 392



scelio Antoine, 1944, Bembidion [Peryphus] 245

scerisorae Knirsch, 1913, Duvalius [Anophthalmus] 300

schachensis Mandl, 1955, Carabus 137

schaeferi J. Jacquet, 1943, Amara [Deletum] 565

schaefleini Brancucci, 1988, Platambus 44



schalleri Gmelin, 1790, Copelatus [Dytiscus] 47

schamsiensis Poppius, 1908, Amara 548

schamsiensis Poppius, 1908, Poecilus [Pterostichus] 485

schamyli Hampe, 1852, Carabus 201

scharifi Morvan, 1977, Ophonus [Metophonus] 391



scharifi Morvan, 1976, Trechus 329

schartowii Heer, 1837, Carabus 171

schaschli Marseul, 1880, Pterostichus [Feronia] 507

schatzmayri J. Müller, 1926, Abax 469

schatzmayri P. Moravec & Lompe, nom. nov., Anophthalmus 22, 292



schatzmayri Born, 1912, Carabus 196

schatzmayri J. Müller, 1912, Duvalius [Trechus] 307

schatzmayri Jeanne, 1981, Poecilus 483

schatzmayri Bari, 1950, Reicheiodes [Dyschirius] 230

schatzmayri Monzini & Pesarini, 1986, Stomis 518

schatzmayri Straneo, 1935, Tapinopterus [Pterostichus] 519

schatzmayri Focarile, 1949, Trechus 22, 326

schaubergeri Jaeger, 1995, Bradycellus 401

schaubergeri Jedlika, 1931, Chydaeus 362

schaubergeri Jedlika, 1932, Harpalomimetes [Anisodactylus] 363



schaubergeri Csiki, 1932, Ophonus [Harpalus] 388

schaubergeri Roubal, 1928, Patrobus 285

schaubergeri Jedlika, 1935, Stenolophus [Egadroma] 405

schaubergerianus Puel, 1937, Ophonus [Metophonus] 391

schaufussi Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 542

schaufussi Putzeys, 1870, Trechus 340

schaumei Aubé, 1842, Nebrioporus [Hydroporus] 66

schaumi A. Morawitz, 1862, Carabus 165

schaumi Birthler, 1886, Carabus 173

schaumi Gaubil, 1849, Carabus 167

schaumi Solsky, 1867, Haliplus 31

schaumi Saulcy, 1863, Microtyphlus [Scotodipnus] 240



schaumii F. J. Schmidt, 1860, Anophthalmus 292

schaumii Schaum, 1860, Anophthalmus 292

schaumii Putzeys, 1866, Dyschirius 229

schaumii Wollaston, 1864, Harpalus 370

schaumii Pandellé, 1867, Trechus 340

schaumii Wollaston, 1854, Zargus 440

schawalleri Hieke, 1990, Amara 563

schawalleri Müller‑Motzfeld, 1995, Bembidion 267

schawalleri Deuve, 1990, Broscosoma 235

schawalleri J. Schmidt, 1998, Deltomerodes 281

schawalleri Baehr, 1998, Gunvorita 573

schawalleri Shilenkov, 1998, Nebria 90

schawalleri Deuve & Hodebert, 1991, Trechus 340

scheerpeltzi Mandl, 1955, Carabus 178

scheerpeltzi Winkler, 1936, Trechus 340



scheffeli Born, 1911, Carabus 185

scheibei Eidam, 1937, Carabus 162

scheibeli Winkler, 1927, Duvalius 303

scheidleri Panzer, 1799, Carabus 173

schelkovnikovi Zaitzev, 1918, Calosoma 119

schelkovnikovi Zaitzev, 1913, Hydaticus 54

schelkownikovi Carret, 1898, Parazuphium [Zuphium] 574

schencki Géhin, 1885, Carabus 184

schenklingi G. Hauser, 1921, Carabus [Coptolabrus] 143


schenklingi Dupuis, 1912, Peripristus [Thyreopterus] 437

schereri Naviaux, 1994, Neocollyris 116

schereri Jedlika, 1964, Peliocypas [Risophilus] 434

schermanni Kirschenhofer, 1985, Bembidion 257

schestopalovi Plutenko, 1995, Carabus 174

scheubeli Mandl, 1955, Carabus 153

scheuerni Baehr, 1982, Atranopsis 521

schildknechti Dettner, 1983, Colymbetes 45

schilenkovi Hieke, 1988, Amara 551

schillhammeri Toledano, 1998, Bembidion 252

schillhammeri Mazzoldi, 1998, Orectochilus 29

schillhammeri Wewalka & Brancucci, 1995, Platambus 44

schilskyi Kraatz, 1887, Carabus 137

schimperi Wencker, 1866, Amara 551

schiodtii Westwood, 1874 Paussus [add] 210

schirasensis Breuning, 1968, Carabus 169

schirmeri Chaudoir, 1859, Laemostenus [Sphodrus] 538

schlegelmilchi M. F. Adams, 1817, Nebria [Carabus] 87

schmalfussi Baehr, 1983, Trechus 340

schmidi Breuning, 1973, Carabus 176

schmidtgoebeli Lorenz, 1998, Lebia 429

schmidtgoebeli W. Horn, 1893, Neocollyris [Collyris] 116

schmidti Zaitzev, 1913, Agabus 36

schmidti Hieke, 1994, Amara 553

schmidti Kirschenhofer, 1991, Apristus 431

schmidti Wollaston, 1854, Bembidion 253

schmidti Andrewes, 1927, Brachinus 216



schmidti Apfelbeck, 1890, Carabus 163

schmidti Heinz, 1994, Cychropsis 202

schmidti Chaudoir, 1837, Cychrus 205

schmidti Zamotajlov, 1999, Deltomerodes 281

schmidti W. Horn, 1927, Hypaetha [Cicindela] 112

schmidti Sciaky & Facchini, 1997, Microzargus 440



schmidti Tschitschérine, 1895, Tachyura [Tachys] 279

schmidtii Sturm, 1844, Anophthalmus 292



schmidtii Chaudoir, 1859, Euplynes 458

schmidtii L. Miller, 1854, Laemostenus [Sphodrus] 535

schmidtii Chaudoir, 1838, Pterostichus [Steropus] 511

schmiedeknechti Pic, 1900, Microlestes 422

schneideri Putzeys, 1878, Amara 565

schneideri Reitter, 1888, Carabus [Neoplectes] 133

schneideri Imura, 1997, Cychrus 205

schneideri Crotch, 1871, Dromius 420

schneideri Ganglbauer, 1915, Zabrus [Eutroctes] 569

schnelli Lassalle, 1999, Carabus 183



schnitteri Jaeger, 1999, Acupalpus 399

schodaicus Kirschenhofer, 1987, Pterostichus 505

schoedli Fery, Fresneda & Millán, 1996, Nebrioporus 67

schoenemanni Kirschenhofer, 1994, Merizomena [Glycia] 409

schoenherri Fischer von Waldheim, 1820, Carabus 164

schoenherri Dejean, 1831, Chlaenius 354

schoenherri Aubé, 1838, Hygrotus [Hydroporus] 72

schoenherri Faldermann, 1836, Pterostichus 501



schoeni Novotný & Voíšek, 1988, Carabus 200

schoenmanni Fischhuber, 1984, Duvalius 304

schoenmanni Wewalka, 1997, Microdytes 75

schoenmanni Kirschenhofer, 1991, Pterostichus 506

schoenmanni Sciaky, 1998, Stomis 518



scholzi W. Kolbe, 1916, Agabus 35

scholzi W. Kolbe, 1899, Hydroporus 63

scholzi Machado, 1992, Paradromius 424

schotti Obydov, 1996, Carabus 160

schouberti Tschitschérine, 1898, Harpalus 374



schoutedeni Burgeon, 1935, Tachys 275

schrammi Antoine, 1941, Amara 555



schrammi Antoine, 1950, Cicindela 103

schrammi Antoine, 1938, Laemostenus 536

schrammi Antoine, 1941, Syntomus [Metabletus] 433

schrankii Duftschmid, 1812, Chlaenius [Carabus] 350

schreiberi Kraatz, 1877, Carabus 163



schreibersii Duftschmid, 1812, Harpalus [Carabus] 375

schreibersii Kuster, 1846, Laemostenus [Sphodrus] 535



schreibersii Dejean, 1831, Nebria 91

schrencki Motschulsky, 1860, Carabus 123

schrenckii Gebler, 1843, Nebria 89

schrenckii A. Morawitz, 1862, Pterostichus 497

schrenki Fischer von Waldheim, 1844, Cicindela 106

schrenkii Gebler, 1841, Cephalota [Cicindela] 102



schrenkii Gebler, 1845, Taphoxenus [Sphodrus] 544

schrickelli Dejean, 1826, Carabus 136

schroederi Holdhaus, 1912, Microlestes 423

schtschukini Kurnakov, 1961, Lindrothius 529

schuberti Jedlika, 1963, Bembidion 255

schuberti Breuning, 1968, Carabus 185



schuberti Breuning, 1971, Carabus 193

schuberti Wewalka, 1971, Deronectes 59



schuberti Jedlika, 1968, Leistus 81

schuberti Jedlika, 1968, Licinus 442

schuberti Jedlika, 1963, Platyderus 523

schuberti Jedlika, 1968, Tachyura [Tachys] 280



schuberti Mandl, 1936, Tapinopterus 519

schubertianus Breuning & Ruspoli, 1974, Carabus 148

schuelkei Jaeger & Wrase, 1996, Bradycellus 402

schuelkei Sciaky & Wrase, 1997, Pterostichus 488

schuelkei P. Moravec & Wrase, 1998, Trechus 340

schueppeli Palliardi, 1825, Abax 469

schueppelii Dejean, 1831, Bembidion 267

schueppelii Dejean, 1825, Cymbionotum [Siagona] 235

schuetzei Kleinfeld, 1998, Carabus 127

schuetzei Sciaky & Wrase, 1997, Myas [Trigonognatha] 476



schuleri J. Jacquet, 1936, Nebria 86

schuleri Jeannel, 1928, Neotrechus 315

schuleri Aubry, 1956, Pterostichus [Haptoderus] 489

schulerianum Müller‑Motzfeld, 1986, Bembidion 259



schurmanni Breuning, 1977, Calosoma 121

schurmanni Machado, 1987, Licinopsis 540

schusteri Ganglbauer, 1889, Oreonebria [Nebria] 96

schusteri Ganglbauer, 1891, Trechus 330

schweigeri Hieke, 1997, Amara 563

schweigeri Korge, 1971, Speluncarius 516

schweigerinae Schweiger, 1962, Carabus 193

schweinfurthi A. Zimmermann, 1921, Nebrioporus [Deronectes] 67

schwienbacheri Grottolo & Martinelli, 1991, Orotrechus 317

sciakyi Toledano, 1999, Bembidion 245

sciakyi J. Schmidt, 1996, Deltomerodes 282

sciakyi Zamotajlov, 1996, Diplous 283

sciakyi Kirschenhofer, 1995, Holosoma 444

sciakyi Casale & Vigna Taglianti, 1999, Laemostenus 538

sciakyi Farka, 1995, Leistus 81

sciakyi Zamotajlov, 1999, Naxipenetretus 284

sciakyi Ledoux & Roux, 1996, Nebria 89

sciakyi Wrase, 1990, Ophonus 391

sciakyi Jeanne, 1996, Platyderus 523



scialdonei G. Hauser, 1913, Carabus [Coptolabrus] 143

scicula Baudi di Selve, 1864, Clivina 220

scintillans Boheman, 1858, Agonum [Anchomenus] 454

scintillans Bates, 1892, Catascopus 435

scintillus Reitter, 1884, Carabus 194



scipio Reitter, 1900, Harpalus 376

scitulum Dejean, 1828, Agonum 453



scitulus Stephens, 1828, Stictonectes [Hygrotus] 69

scitus Mäklin, 1878, Pterostichus [Feronia] 492

scitus Jeannel, 1960, Trechus 340

sclanoi Magrini, 1996, Bembidion [Ocydromus] 245

sclanoi Magrini & Vanni, 1996, Duvalius 305


sclopeta Fabricius, 1792, Brachinus [Carabus] 214

scolecaster Gistel, 1857, Carabus 171

scopolii Gmelin, 1790, Acilius [Dytiscus] 49

scopolii F. J. Schmidt, 1850, Anophthalmus 293

scopolii Sturm, 1851, Anophthalmus 293

scopolii Dejean, 1826, Carabus 180

scopularis Schiødte, 1841, Hydroporus 64

scopulinum Kirby, 1837, Bembidion [Peryphus] 255



scordiscus Lapouge, 1901, Carabus 166

scorteccii Schatzmayr, 1940, Bembidion 259

scoteinus Kolenati, 1845, Brachinus 213

scoticus Stephens, 1828, Acilius 49

scotomedes L. Redtenbacher, 1867, Brachinus 216

scotti Basilewsky, 1948, Afrotarus [Cymindis] 411

scotti Liebke, 1934, Brachinus 216



scotus Jeannel, 1942, Pterostichus [Platysma] 509

scotus Habu, 1977, Takasagoagonum 467

scovitzi Faldermann, 1836, Carabus 194

scripta Putzeys, 1846, Clivina 220

scripta Clairville, 1806, Cylindera [Cicindela] 110

scriptifrons A. Fleischer, 1898, Dyschirius 227



scriptus Régimbart, 1895, Metaporus [Hydroporus] 65

scrobiculatus Fischer von Waldheim, 1822, Carabus 147

scrobiculatus Fabricius, 1801, Platynus [Carabus] 463



scrutata Casey, 1918, Amara [Curtonotus] 561

scrutatum Casey, 1918, Bembidion 271

sculpticollis Reitter, 1887, Poecilus [Pedius] 480

sculptilis Heer, 1837, Carabus 185

sculptilis Putzeys, 1866, Clivina 220

sculptior Deuve, 1992, Carabus 147

sculptipennis Chaudoir, 1877, Carabus 198

sculptipennis Sirguey, 1931, Carabus 139

sculptipes Bates, 1883, Agonum [Anchomenus] 451

sculpturatus Ménétriés, 1832, Carabus 137

sculpturatus Régimbart, 1884, Orectochilus 29



sculptus Lapouge, 1925, Carabus 161


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