Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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baeterrensis Lapouge, 1901, Carabus 165

baetica Rambur, 1837, Cymindis 413

baeticum K. Daniel & J. Daniel, 1898, Parazuphium [Zuphium] 573

baeticus Rambur, 1837, Calathus 524

baeticus Rambur, 1837, Chlaenius 352

baeticus Rambur, 1837, Eocarterus [Ditomus] 365

baeticus Rambur, 1837, Laemostenus [Pristonychus] 537

baeticus Schaum, 1864, Nebrioporus [Hydroporus] 67

baeticus Nègre, 1958, Paradromius [Dromius] 425

baeticus Rambur, 1838, Poecilus 479

bagdadensis Liebke, 1928, Lebia 428

bagdadica Tschitschérine, 1898, Amara 567

bagdatensis Pic, 1902, Brachinus 214

baghlanicum Kirschenhofer, 1989, Bembidion 256

bagrovdaghensis Shilenkov, 1983, Nebria 85

baguenai Breuning, 1926, Carabus [Mesocarabus] 167

bahreinica Ali, 1978, Calomera [Cicindela] 99

baiburtensis Breuning, 1975, Cychrus 202



baicalensis Semenov, 1895, Cymindis 415

baicalensis Motschulsky, 1844, Dyschirius 227



baicalensis A. Morawitz, 1862, Nebria 88

baicalica Motschulsky, 1844, Nebria 88

baicalicum Motschulsky, 1844, Bembidion [Campa] 263

baidulyensis Belousov & Kabak, 1994, Carabus 150

baikaloussuricum Netolitzky, 1939, Bembidion 247

baima Farka, 1999, Leistus 80

baimanorum Deuve, 1999, Carabus 128

bajamarensis Freude, 1986, Zabrus 570

bajani Jedlika, 1964, Amara 554

bajani Jedlika, 1966, Bembidion 243

bajankoli Belousov & Kabak, 1992, Trechus 327

bajaurae Andrewes, 1924, Stenolophus 405

bajeti J. Vives & E. Vives, 1989, Acinopus 368



bakalowiczi Coiffait, 1973, Duvalius [Trechopsis] 303

bakeri Peschet, 1921, Canthydrus 33

bakurovi Shilenkov, 1984, Trechus 327

balachowskyi Morvan, 1970, Carabus 138

balachowskyi Ledoux, 1976, Deltomerus 282



balachowskyi Morvan, 1970, Laemostenus [Antisphodrus] 534

balachowskyi Morvan, 1972, Pterostichus 509



balanensis Csiki, 1942, Carabus 167

balangensis Xie & Yu, 1993, Aristochroa 470

balangensis Zamotajlov & Sciaky, 1996, Parapenetretus 284

balangshana Hieke, 1994, Amara 553

balassogloi C. A. Dohrn, 1882, Carabus 145


balassogloi Tschitschérine, 1887, Poecilus [Sogines] 481

balazuci Fongond, 1982, Amara [Paracelia] 565

balazuci Bruneau de Miré, 1948, Duvalius 301



balbii Bonelli, 1810, Nebria 88

balcanica Baliani, 1939, Amara 558

balcanicum Netolitzky, 1918, Asaphidion 242

balcanicum Apfelbeck, 1899, Bembidion 256

balcanicus Hlisnikovský, 1955, Agabus 38

balcanicus Born, 1899, Carabus 195

balcanicus Lapouge, 1901, Carabus 166

balcanicus Hopffgarten, 1881, Cychrus 205

balcanicus J. Frivaldszky, 1879, Duvalius [Trechus] 308

balcanicus Mlyná, 1977, Molops 473

balcanicus Ganglbauer, 1891, Tapinopterus 519

balcanicus Jeannel, 1927, Trechus 23, 332

balcanicus Heyden, 1883, Zabrus 570

balchashica Emetz & Kryzhanovskij, 1973, Cymindis 415

baldense Rosenhauer, 1846, Broscosoma 235



baldensis Depoli, 1939, Abax 470

baldensis K. Daniel & J. Daniel, 1898, Amara 563

baldensis Schaum, 1856, Carabus 184

baldensis Putzeys, 1870, Duvalius [Trechus] 301



baldensis Jeannel, 1937, Laemostenus [Cryptotrichus] 536

baldensis Schaum, 1862, Pterostichus 488

balearica Sydow, 1934, Cicindela 103

balearica Español, 1974, Reicheia 222



balearicus Lapouge, 1913, Carabus 161

balearicus Henrot, 1964, Duvalius 301

balearicus Piochard de la Brûlerie, 1868, Orthomus [Feronia] 476

balearicus Jeanne, 1970, Platyderus 522

balfourbrownei Vazirani, 1965, Platambus 43

balfourbrownei Uéno, 1965, Trechus 327

baliani Jedlika, 1935, Amara 567

balianica Fassati, 1952, Amara 560

balkaricus Belousov & Abdurachmanov, 1991, Carabus 200

balkaricus Belousov & Komarov, 1998, Nannotrechus 314

balkaricus Belousov, 1990, Trechus 327

balkei Fery & Hosseinie, 1998, Deronectes 58

balkei Fery & Nilsson, 1993, Ilybius [Agabus] 41

balkenohli Sciaky, 1996, Laemostenus 535

balkhashica Kabak, 1993, Amara [Cribramara] 558



ballassogloi Dokhtouroff, 1882, Cicindela 105

balli Sciaky & Facchini, 1999, Loricera 99

balli Vazirani, 1970, Nebrioporus [Potamonectes] 66

balli Uéno, 1986, Trechiama 322



ballioni Solsky, 1874, Calosoma [Callisthenes] 118

ballioni Heyden, 1882, Harpalus 373

ballioni Kirschenhofer, 1990, Harpalus 382

ballionis Kraatz, 1879, Carabus 178



ballionis Retowski, 1888, Cychrus 202

ballionis Heyden, 1880, Mnuphorus [Lebia] 358

ballista Germar, 1823, Aptinus 212

balmae Delarouzée, 1860, Laemostenus [Pristonychus] 533

balsetensis Abeille de Perrin, 1904, Siettitia 69

baltalensis Ledoux, 1980, Carabus 155

balteata Heyden, 1886, Lebia 428



balthasari Jedlika, 1951, Bembidion 265

balthasari Jedlika, 1940, Platynus [Colpodes] 464

balthasari Jedlika, 1937, Pterostichus 488

baltica Motschulsky, 1844, Cicindela 106

baltica Motschulsky, 1866, Nebria 88

baltimorensis Herbst, 1806, Cylindera [Cicindela] 111

balucha Bates, 1878, Cylindera [Cicindela] 111

bambousicola Deuve & Tian, 1999, Carabus 128

bameuli Fery & Hosseinie, 1998, Deronectes 58

bamidunyae Bates, 1878, Amara 548

banajai Brancucci, 1980, Nebrioporus [Potamonectes] 66

banaticus L. Redtenbacher, 1849, Carabus 186

banaticus I. Frivaldszky von Frivald, 1865, Platynus [Anchomenus] 462

bandoi Uéno, 1986, Trechiama 322

bandotaro Morita, 1991, Bembidion 271

bandotaro Tanaka, 1958, Pterostichus 486

bangalorensis Vazirani, 1970, Copelatus 47

banghaasi Baliani, 1933, Amara 559

banghaasi Reitter, 1895, Carabus 153

banksi Crotch, 1871, Nebria 89



banksii Crotch, 1872, Hydaticus 54

bannaticus Dejean, 1828, Pterostichus [Cophosus] [ND] 515

bannaticus Dejean, 1831, Trechus 327

banoni Dejean, 1829, Carabus 186

baojiensis Imura, 1993, Cychrus 203

baoshanensis Deuve, 1993, Carabus 127

barahbisensis Deuve, 1988, Trechus 327

barajoni Focarile, 1949, Trechus 340

barakaicus Zamotajlov, 1991, Carabus 200



baranyensis Sokolá, 1908, Carabus 152

baratellii Sciaky, 1985, Anophthalmus 291



barbara Laporte, 1840, Calomera [Cicindela] 101

barbara Chaudoir, 1843, Nebria 92

barbarae Sciaky & Facchini, 1999, Loricera 98



barbarum J. Müller, 1918, Bembidion 267

barbarus Bedel, 1904, Aephnidius 357

barbarus Lucas, 1846, Brachinus 212

barbarus Dejean, 1826, Carabus 161

barbarus Solier, 1835, Carterus [Odogenius] 363

barbarus Schaum, 1847, Herophydrus [Hydroporus] 71

barbarus Lucas, 1846, Laemostenus [Pristonychus] 536



barbarus Leder, 1872, Ophonus [Dichirotrichus] 391

barbarus Dejean, 1828, Orthomus [Feronia] 477



barbarus Lucas, 1842, Poecilus 483

barbarus Puel, 1938, Syntomus [Metabletus] 432

barbata Andrewes, 1929, Nebria 90

barbatus Wollaston, 1862, Calathus 526

barbatus Fischer von Waldheim, 1829, Pterostichus [Poecilus] [ND] 515

barberae Ortuño, 1991, Perileptus 289

barbimentosa Shilenkov, 1983, Nebria 85

barbulatus Cavazzuti & Ratti, 1999, Carabus 128

barcelecoanus Lapouge, 1925, Carabus 167

barcelonensis Hieke, 1983, Amara 554

barcelonensis Schulz, 1909, Cylindera [Cicindela] 110

baretosanus Jeannel, 1941, Aphoenops 295

bargusinica Shilenkov, 1999, Nebria 88

bargusinicus Shilenkov, 1996, Carabus 172

barii Focarile, 1949, Trechus 327

barkamense Toledano, 1998, Bembidion 251

barkamensis Deuve, 1998, Amerizus 241

barkamensis Deuve, 1994, Carabus 191

barkamensis Deuve, 1991, Cychrus 203

barkamensis Farka, 1995, Leistus 80



barkamianus Korell & Kleinfeld, 1988, Carabus 154

barkamicola Deuve, 1996, Carabus 189

barlensis Straneo, 1935, Pterostichus 489

barmaticus Saha, 1984, Chlaenius 354

barnevillei Croissandeau, 1893, Amara 568

barnevillei Fairmaire, 1856, Amara 557

barnevillei Chaudoir, 1867, Leistus 81

barnevillei Pandellé, 1867, Trechus 327

barnolai Xaxars, 1921, Carabus 161

baronii Heinertz, 1977, Carabus 154

barovskii Semenov & Znojko, 1932, Carabus 123

barrancoi Mateu, 1997, Laemostenus 533

barratxinai Español, 1971, Trechus 327

barrosi Lapouge, 1908, Carabus 130

barrosi Jeanne, 1996, Platyderus 522



barthei Puel, 1939, Calathus 527

barthei Lapouge, 1912, Carabus 139

barthei Puel, 1937, Carterus 364

barthei Antoine, 1920, Harpalus 380

barthei Puel, 1923, Pogonus 286

barthei Jeannel, 1947, Pterostichus [Haptoderus] 489



barthei Jeannel, 1949, Pterostichus [Argutor] [HN] 507

barthelemyi Gené, 1836, Calomera [Cicindela] 101

barthelemyi Gory, 1838, Cephalota [Cicindela] 102

barthelemyi Dejean, 1830, Graphipterus 359



bartholomaei Motschulsky, 1852, Carabus 160

bartkoi Csiki, 1912, Anophthalmus 293

bartoni Maan, 1930, Carabus 165

bartoni Maan, 1934, Licinus 442

bartoni Maan, 1935, Ophonus [Harpalus] 388

bartonii Maan, 1933, Tapinopterus 519

bartschi Wrase, 1998, Bradycellus 401

barysomus Bates, 1889, Carabus 154



basale Puel, 1938, Agonum 451

basale Chaudoir, 1846, Bembidion 261

basale L. Miller, 1868, Bembidion 268

basale Motschulsky, 1844, Bembidion [Peryphus] 20, 249

basalis Gebler, 1829, Agabus [Colymbetes] 38

basalis Newman, 1838, Broscus 236

basalis Schilsky, 1888, Calodromius [Dromius] 419

basalis Chaudoir, 1872, Catascopus 435

basalis Fischer von Waldheim, 1828, Chlaenius 350

basalis Boheman, 1848, Copelatus 48

basalis Gyllenhal, 1810, Cymindis 418

basalis A. Fleischer, 1899, Dyschirius 225

basalis Motschulsky, 1844, Harpalus 386

basalis Sharp, 1882, Lacconectus 48

basalis Motschulsky, 1859, Laccophilus 76

basalis Chaudoir, 1852, Lebia [Cymindis] 429

basalis Chaudoir, 1852, Merizomena [Lebia] 409

basanicus J. R. Sahlberg, 1908, Anthracus 399

basanicus J. R. Sahlberg, 1913, Harpalus 373

basarukini P. Moravec & Wrase 1997, Trechus 327



baschkirica Schweiger, 1966, Amara [Bradytus] 553

baschkiricus Breuning, 1932, Carabus 135



baschkiricus Motschulsky, 1844, Elaphrus 207

bashanensis Imura, 1995, Carabus 128

basharicus Schauberger, 1933, Harpalus 375

basiclarior Gozis, 1914, Hydroporus 62

basicollis Fairmaire, 1892, Idiomelas [Anisodactylus] 404

basilaris Kirby, 1837, Harpalus 376


basilevskyi Lutshnik, 1935, Amara 558

basilewskianus Breuning & Ruspoli, 1970, Carabus 194

basilewskyi Breuning, 1966, Carabus 201

basilewskyi Kult, 1954, Dyschirius 228

basilewskyi Luna de Carvalho, 1967, Protopaussus 19, 211

basilewskyi Casale, 1988, Pseudotaphoxenus 540

basilianus Stark, 1890, Carabus 132



basilicatus Born, 1902, Carabus [Procrustes] 187

basilicus Chevrolat, 1836, Carabus 140

basinotatus A. Zimmermann, 1924, Haliplus 31

basinotatus Reiche, 1864, Hydroporus 60

basipes Reitter, 1887, Nebria 87

basiplicatus Heyden, 1884, Scarites 233

basipunctatus Straneo, 1955, Pterostichus 511

basirufum Holdhaus, 1910, Bembidion 256

basisquamatus Uéno, 1982, Yamautidius 345

baskonicus Belousov & Kabak, 1996, Trechus 327

basoglui Schweiger, 1964, Carabus 185

basquini Darge, 1990, Carabus 193

basquinianus Darge, 1992, Carabus 158

basrensis Ali, 1967, Oodes 445



bassieri A. Gaudin, 1947, Aphoenops 295

bastiani Lassalle, 1983, Carabus 130

bastianinii Magrini, 1998, Duvalius 301

bastianinii Magrini & Vanni, 1994, Hypotyphlus 239

batangensis Deuve, 1989, Carabus 175

batangicus Deuve, 1989, Carabus 190

batesi Csiki, 1929, Amara 553

batesi Putzeys, 1875, Bembidion [Notaphus] 253

batesi Semenov, 1891, Broscus 236

batesi Lapouge, 1916, Carabus 154

batesi Fleutiaux, 1894, Cosmodela [Cicindela] 108

batesi Habu, 1958, Dromius 420

batesi Andrewes, 1926, Dyschirius 229

batesi Harold, 1877, Euplynes 458



batesi Lewis, 1896, Eustra 208

batesi Kanô, 1929, Lebidia 426

batesi Tschitschérine, 1894, Pterostichus [Feronia] 500

batesi Chaudoir, 1876, Styphlomerus 218

batesianus Deuve, 1995, Carabus 172

batesianus Lutshnik, 1916, Poecilus [Platysma] 481

batesoni Semenov & Redikorzev, 1928, Calosoma 119

batnense Lallemant, 1868, Calosoma 119

batschkowskii B. Bodemeyer, 1928, Carabus [Plectes] 132

battalensis Heinz & Staven, 1998, Carabus 154



battoniellus Deuve, 1991, Carabus 175

battoniensis Deuve, 1991, Carabus 128

battonii Straneo, 1959, Duvalius 301

battonii Straneo, 1959, Pterostichus 511

battus Reitter, 1900, Ophonus 391

batyr Belousov & Kabak, 1992, Trechus 327

baudii A. Fiori, 1913, Acinopus 368

baudii Seidlitz, 1887, Agabus 40

baudii Kraatz, 1878, Carabus 180

baudii Fairmaire, 1892, Microlestes 422



baudii Ganglbauer, 1891, Oreonebria [Nebria] 96

baudii Chaudoir, 1868, Pterostichus [Feronia] 503

baudii Ragusa, 1883, Typhloreicheia 223

baudinoti Deuve, 1985, Leistus 80

baudoni Antoine, 1952, Dyschirius 224

baudoni Antoine, 1957, Poecilus 481

baudoni Antoine, 1957, Zabrus 572

baudueri Perris, 1869, Pseudomasoreus [Cymindis] 418

baumi Müller‑Motzfeld, 1985, Bembidion 261

baxiensis Deuve, 1998, Amerizus 241

baxiensis Deuve, 1997, Cychrus 203

bayanbulak Deuve, 1993, Carabus 123

bayanbulak Deuve, 1993, Trechus 327

bayardi Dejean, 1831, Brachinus 214

bayardi Solier, 1835, Carabus 138



bayeri Lapouge, 1910, Carabus 168

baznosanui Mallász, 1928, Duvalius 298



baznosanui Mallász, 1930, Duvalius 298

baztanensis Dupré, 1991, Trechus 327

beatricis Magrini, Cecchi & Lo Cascio, 1998, Ocys 272

beatus Reitter, 1903, Trechus 327



beauchenii Fairmaire, 1889, Mimocolliuris [Ophionea] 444

beauvoisi Dejean, 1826, Carabus 168

beccarii Putzeys, 1873, Coryza 221

beccarii Gestro, 1889, Lionychus 431

bechteli Charet, 1996, Carabus 193

bechynei Kult, 1949, Dyschirius 230

beckenhauptii Duftschmid, 1812, Abax [Carabus] 469

beckeri Jedlika, 1931, Dolichus 530

beckeri Jedlika, 1930, Pheropsophus 217

beckianus Ganglbauer, 1888, Omphreus 446

beckmanni Ahlwarth, 1913, Haliplus 33

becvari Dostal, 1999, Distichus 231

becvari Farka, 1999, Leistus 80

becvari Sciaky, 1995, Myas [Trigonognatha] 475



bedelensis Janák & P. Moravec, 1989, Duvalius 298

bedeli Netolitzky, 1914, Bembidion 270

bedeli Tschitschérine, 1897, Chydaeus [Acrogeniodon] 361



bedeli A. Nicolas, 1906, Cillenus 272

bedeli Tschitschérine, 1897, Cymindis 415



bedeli J. Müller, 1922, Dyschirius 227

bedeli Zaitzev, 1908, Ilybius [Agabus] 42



bedeli Holdhaus, 1912, Microlestes 423

bedeli Peyerimhoff, 1908, Ocys [Oreocys] 272

bedeli Escalera, 1913, Phloeozetus [Singilis] 438

bedeli Antoine, 1957, Platyderus 522

bedeli Puel, 1924, Syntomus [Metabletus] 432

bedeli Jeannel, 1922, Trechus 327

bedeli Vauloger de Beaupré, 1897, Zuphium 574

bedelianum Netolitzky, 1918, Bembidion 270

bedelianus Lutshnik, 1915, Orthomus [Platysma] 476

bedoci Jeannel, 1923, Neoblemus 288

bedosae Deuve, 1996, Sinotroglodytes 320



beesoni Baliani, 1934, Amara 548

beesoni Andrewes, 1933, Bembidion 269



beesoni Andrewes, 1919, Calosoma 119

beesoni Andrewes, 1927, Poecilus [Pterostichus] 481

beesoni Jeannel, 1930, Trechus 327

beethami Andrewes, 1930, Aparupa 456



begeri Jedlika, 1968, Amara 557

beghinorum Belousov & Kabak, 1992, Trechus 327

begoueni Jeannel, 1926, Geotrechus 310

beharrius A. Nicolas, 1916, Carabus 140

behningi Zaitzev, 1927, Graptodytes 60

beicki Hieke, 1988, Amara 559

beidensis Antoine, 1941, Philorhizus 425

beieri Winkler, 1936, Trechus 327



beirensis Born, 1924, Carabus 146

bejarana Lauffer, 1905, Cymindis 411

bejarensis Kirschenhofer, 1979, Pterostichus 497

bejarensis Jeannel, 1927, Trechus 340



belasicensis Mandl, 1989, Cychrus 205

belasicensis Mlyná, 1977, Molops 474

belasicensis Maan, 1933, Tapinopterus 519

belenae Zaballos, 1983, Typhlocharis 241



belgicus Lapouge, 1913, Carabus 168

belizini Lutshnik, 1933, Amara 555

belizini Lutshnik, 1933, Pterostichus [Platysma] 504

bella Habu, 1979, Teradaia 437

bellana W. Horn, 1905, Salpingophora [Cicindela] 114



bellasilviae J. Schmidt, 1996, Xestagonum 467

bellatrix Tschitschérine, 1895, Pterostichus [Feronia] 497

bellesi Lagar Mascaró, 1971, Catalanodytes [Typhloreicheia] 221

belli Andrewes, 1927, Platynus [Anchomenus] 463

belli Andrewes, 1925, Tachyura [Tachys] 278

bellicosus L. Dufour, 1820, Brachinus 212

bellicum Lutshnik, 1934, Agonum 452

bellieri Gautier des Cottes, 1867, Calathus 528

bellieri Reiche, 1861, Harpalus 373

bellieri Reiche, 1861, Orectochilus 28

belloni Tarrier, 1965, Carabus 140

belloti Franz, 1954, Nebria 92



bellulum Casey, 1918, Bembidion 255

bellum C. R. Sahlberg, 1827, Bembidion 244

bellus Kraatz, 1886, Carabus 135

bellus Habu, 1978, Synuchus 545

beloborodovi Belousov & Mikhailov, 1990, Bembidion 245

beloni Raffray, 1870, Laemostenus [Pristonychus] 536

belousovi Kabak, 1992, Carabus 147



belousovi O. Berlov, 1997, Cychrus 19, 202

belousovi Zamotajlov, 1988, Deltomerus 282

belousovi Kryzhanovskij, 1989, Pterostichus 504

belousovi P. Moravec & Wrase, 1995, Trechus [Epaphius] 23, 325

belovi Belousov & Kabak, 1996, Trechus 327

beltrani Toribio, 1990, Trechus 327

beludscha Tschitschérine, 1904, Dyschirius 227

bembidioides Bates, 1889, Arhytinus 456

bembidioides Jedlika, 1936, Diacanthostylus [Colpodes] 458

benabarrensis Pierre Meyer & Mollard, 1996, Carabus 168


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