References
Altringham, J.D., O'Brien, S. & Julian, S. (1994) Feeding ecology of the little owl (Athene
noctua) an an upland site in northern England. Naturalist 119: 81-94.
Bevan, G. (1964) The food of tawny owls in London. London Bird Report 29: 56-72. Brown, R., Ferguson, J., Lawrence, M. & Lees, D. (1987) Tracks and Signs of the Birds of Britain and Europe. Christopher Helm, London.
Macdonald,A.W.(1976) Nocturnal observations of tawny owls (Strix aluco) preying upon
earthworms. Ibis 118: 579-80.
Mead, C. (1987) Owls. Whittet, London.
Southern, H.N. (1954) Tawny owls and their prey. Ibis 96: 384-410.
Wardhaugh, A.A. (1983) Owls of Britain and Europe. Blandford, Poole.
Yalden, D.W. (1977) The Identification of Remains in Owl Pellets. The Mammal Society,London,
Table 1:Monthly Record of Pellets Collected and Presence of Owls at Roost
Collection Date
|
No. of Pellets
|
Volume (cm3)
|
No. of Owls Seen
|
|
|
|
|
24. 1. 93
|
17
|
-
|
1
|
19. 2. 93
|
12
|
-
|
2
|
27. 3. 93
|
16
|
127
|
0
|
21. 4. 93
|
20
|
138
|
0
|
22. 5. 93
|
14
|
77
|
0
|
21. 6. 93
|
7
|
39
|
0
|
22. 7. 93
|
16
|
104
|
0
|
22. 8.93
|
0
|
0
|
0
|
25. 9. 93
|
1
|
5
|
0
|
25.10.93
|
1
|
4
|
2
|
20.11. 93
|
6
|
38
|
2
|
20.12. 93
|
14
|
82
|
2
|
22. 1. 94
|
7
|
29
|
1
|
20. 2. 94
|
6
|
30
|
0
|
26. 3. 94
|
32
|
187
|
1
|
24. 4.94
|
15
|
103
|
0
|
23. 5. 94
|
1
|
6
|
0
|
22. 6.94
|
1
|
3
|
0
|
25. 7.94
|
1
|
6
|
0
|
22. 8. 94
|
0
|
0
|
0
|
25. 9.94
|
0
|
0
|
0
|
26.10.94
|
0
|
0
|
0
|
19.11.94
|
0
|
0
|
0
|
22.12. 94
|
1
|
6
|
0
|
TOTAL
|
159
|
984
|
-
|
Table 2:Monthly Record of Prey Items
Collection Date
|
Bank Vole
|
Short-tailed Vole
|
Wood Mouse
|
Common Shrew
|
Pygmy Shrew
|
Other
|
24. 1. 93
|
26
|
2
|
18
|
0
|
1
|
1 bird
|
19. 2.93
|
12
|
0
|
5
|
0
|
0
|
|
27. 3. 93
|
14
|
4
|
8
|
1
|
0
|
|
21. 4. 93
|
9
|
5
|
4
|
2
|
1
|
1 probable rat
|
22. 5.93
|
4
|
5
|
2
|
3
|
0
|
|
21. 6.93
|
2
|
3
|
2
|
0
|
0
|
|
22. 7.93
|
9
|
2
|
3
|
2
|
0
|
|
22. 8.93
|
0
|
0
|
0
|
0
|
0
|
|
25. 9. 93
|
1
|
0
|
0
|
0
|
0
|
|
25.10.93
|
1
|
0
|
0
|
0
|
0
|
1bird
|
20.11. 93
|
7
|
2
|
0
|
0
|
0
|
|
20.12. 93
|
7
|
1
|
8
|
2
|
0
|
|
22. 1.94
|
4
|
0
|
1
|
0
|
0
|
|
20. 2. 94
|
3
|
2
|
1
|
0
|
0
|
|
26. 3. 94
|
4
|
11
|
6
|
2
|
4
|
1 bird (robin?)
|
|
|
|
|
|
|
1 young rabbit
|
24. 4. 94
|
4
|
6
|
1
|
2
|
0
|
1 probable
|
|
|
|
|
|
|
young rabbit*
|
23. 5.94
|
0
|
0
|
0
|
0
|
0
|
|
22. 6. 94
|
0
|
1
|
0
|
0
|
0
|
|
25. 7.94
|
1
|
0
|
0
|
0
|
0
|
|
22. 8.94
|
0
|
0
|
0
|
0
|
0
|
|
25. 9.94
|
0
|
0
|
0
|
0
|
0
|
|
26.10. 94
|
0
|
0
|
0
|
0
|
0
|
|
19.11.94
|
0
|
0
|
0
|
0
|
0
|
|
22.12.94
|
2
|
0
|
0
|
0
|
0
|
|
TOTAL
|
111
|
43
|
59
|
14
|
6
|
rat 1, rabbit 2
|
|
|
|
|
|
|
mole 2, bird 3
|
NO. PER100 cm3 **
|
7.42
|
4.17
|
3.66
|
1.42
|
0.61
|
|
*Remains of rabbits found in pellets collected on 26.3.94 and 24.4.94 could be from the
same individual but here counted as two.
** For March 1993 to December 1994 inclusive. (Pellet volumes not recorded January and February 1993).
Total number of prey items identified: 241
Number of prey items per 100cm3 (all species): 17.99
Table 3: Summary of Prey Items Identified
Species
|
Number of Individuals
|
|
Mass Represented*
|
|
|
(%)
|
g
|
(%)
|
Bank vole
|
111
|
(46.1)
|
1776
|
(43.6)
|
Short-tailed vole
|
43
|
(17.8)
|
903
|
(22.1)
|
Wood mouse
|
59
|
(24.5)
|
1062
|
(26.0)
|
Common shrew
|
14
|
(5.8)
|
112
|
(2.7)
|
Pygmy shrew
|
6
|
(2.5)
|
24
|
(0.6)
|
Mole
|
4
|
(0.8)
|
140
|
(3.4)
|
Probable Rat
|
1
|
(0.4)
|
60
|
(1.4)
|
Probable Rabbit
|
2
|
(0.8)
|
-
|
-
|
Bird
|
3
|
(1.2)
|
-
|
-
|
|
|
|
|
|
Total
|
241
|
4077
|
-
|
-
|
* Based on figures for mean mass of each prey species provided in Yalden (1977)
For rabbit and birds there were insufficient remains to estimate size and therefore mass
hence these species were not included in the calculation of percentage mass.
Table 4:Prey Items Identified in Tawny Owl Pellets from elsewhere in the Cleveland Area
Site
|
Date
|
Prey Items
|
Comments
|
|
|
|
|
Cross Wood, Flatts Lane
(NZ 543169)
|
Spring
1967
|
short-tailed vole 1
bank vole 3
bird 1
|
Roost site approx. 0.6 km NW of site of present study; beech tree cavity. Unrecorded small no. of pellets.
|
|
|
|
|
Wood at Ormesby Hall
(NZ 530171)
|
5.4.1971
|
wood mouse 4
common shrew 1
house sparrow 10
|
Roost in a yew tree. Unrecorded small no.of pellets.
|
|
|
|
|
Plantation near incline, Ingleby Greenhow
(NZ 602025)
|
2.4.1978
|
short-tailed vole 2
bird 5
|
Roost in a holly tree. Unrecorded small no. of pellets.
|
|
|
|
|
Wood at Teesside High School
(NZ 425149)
|
9. 6.93
|
birds 2
|
48 cm3 of pellets
|
|
5. 7.93
|
birds 4
|
50cm3 of pellets
|
|
6.12.93
|
wood mouse 4
|
50 cm3 of pellets
|
|
|
brown rat 2
|
Pellets collected from a variety of sites within the wood
|
.
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