Y-dna haplogroup r and its Subclades 2014



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Y-DNA Haplogroup R and its Subclades - 2014

The entire work is identified by the Version Number and date given on the Main Page.   Directions for citing the document are given at the bottom of the Main Page.


Version History     Last revision date for this specific page: 18 October 2014

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Alice Fairhurst if the differences need clarification or if you find any broken links on this page.



LINKS:  Main Page   Y-DNA Tree Trunk   SNP Index   Papers/Presentations Cited   Glossary   Listing Criteria

CLADE/SUBCLADE SYMBOLS:  Added  Redefined 

SNP SYMBOLS:  Not on 2013 tree  Confirmed within subclade  Provisional  Private  Investigation 

The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing one in multiple labs, and/or being found in the area of the chromosome used in recent research studies.

SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup.



Contact people for Haplogroup R:
R1a Contact: Lawrence Mayka                                                     
R1b-U106 and Subclades Contact: Charles Moore                      
R1b-P312 and Subclades Contacts: Alice Fairhurst or Mark Jost
  R1b-P312->DF27 Contact: Stephen Parrish                      
  R1b-P312->L21->DF21 Contact: Charles Fueston           
  R1b-P312->L21->DF49 Contact: David Stedman            
  R1b-P312->L21->CTS4466 Contact: Kathleen Kerwin    
R2 and Subclades Contact: Abdulaziz Ali                                   
Remainder of R Tree Contact: Bob May                                      

Link to Haplogroup R Frequently Mutating SNPs, Private SNPs, Notes and References (Papers)


S18950

S18951

S3262/CST6428
R   M207/Page37/UTY2, P224/PF6050, P227, P229/PF6019, P232, P280, P285, PF6014/S9 M734/PF6057/S4/YSC0000201
R*   -
R1   M173/P241/Page29, M306/PF6147/S1, P225, P231, P233/PF6142, P234, P236, P242/PF6113, P245/PF6117, P286/PF6136, P294/PF6112
• • R1*   -

• • R1a   L62/M513/PF6200, L63/M511/PF6203, L145/M449/PF6175, L146/M420/PF6229


• • • R1a*   -
• • • R1a1   M459/PF6235, L120/M516/PF6236, L122/M448/PF6237, Page65.2/PF6234/SRY1532.2/SRY10831.2
• • • • R1a1*   -
• • • • R1a1a   M512/PF6239, L168, L449/PF6223, M17, M198/PF6238, M514/PF6240, M515
• • • • • R1a1a*   -
• • • • • R1a1a1   M417, Page7
• • • • • • R1a1a1*   -
• • • • • • R1a1a1a   CTS7083/L664/S298
• • • • • • R1a1a1b   S224/Z645, S441/Z647
• • • • • • • R1a1a1b*   -
• • • • • • • R1a1a1b1   PF6217/S339/Z283
• • • • • • • • R1a1a1b1*   -
• • • • • • • • R1a1a1b1a   S198/Z282
• • • • • • • • • R1a1a1b1a*   -
• • • • • • • • • R1a1a1b1a1   M458/PF6241
• • • • • • • • • • R1a1a1b1a1*   -
• • • • • • • • • • R1a1a1b1a1a   L260/S222
• • • • • • • • • • R1a1a1b1a1b   CTS11962
• • • • • • • • • • • R1a1a1b1a1b*   -
• • • • • • • • • • • R1a1a1b1a1b1   L1029/S4554
• • • • • • • • • R1a1a1b1a2   S466/Z280, S204/Z91
• • • • • • • • • • R1a1a1b1a2*   -
• • • • • • • • • • R1a1a1b1a2a   S205/Z92, S344/Z660, S446/Z661
• • • • • • • • • • R1a1a1b1a2b   CTS1211
• • • • • • • • • • • R1a1a1b1a2b*   -
• • • • • • • • • • • R1a1a1b1a2b1   P278.2
• • • • • • • • • • • R1a1a1b1a2b2   L784
• • • • • • • • • • • R1a1a1b1a2b3   CTS3402/S3361
• • • • • • • • • • • • R1a1a1b1a2b3*   -
• • • • • • • • • • • • R1a1a1b1a2b3a   L365/S468
• • • • • • • • • • • • • R1a1a1b1a2b3a*   -
• • • • • • • • • • • • • R1a1a1b1a2b3a1   F2686
• • • • • • • • • • • • R1a1a1b1a2b3b   L366
• • • • • • • • • • • • R1a1a1b1a2b3c   L1280
• • • • • • • • • R1a1a1b1a3   S221/Z284, S443/Z289
• • • • • • • • • • R1a1a1b1a3*   -
• • • • • • • • • • R1a1a1b1a3a   L448/S200
• • • • • • • • • • • R1a1a1b1a3a*   -
• • • • • • • • • • • R1a1a1b1a3a1   CTS4179
• • • • • • • • • • • • R1a1a1b1a3a1*   -
• • • • • • • • • • • • R1a1a1b1a3a1a   L176.1/S179.1
• • • • • • • • • • R1a1a1b1a3b   S223/Z287, S345/Z288
• • • • • • • • • • • R1a1a1b1a3b*   -
• • • • • • • • • • • R1a1a1b1a3b1   CTS8401
• • • • • • • • • • • • R1a1a1b1a3b1*   -
• • • • • • • • • • • • R1a1a1b1a3b1a   S201/Z87, S342/Z88
• • • • • • • • • • • R1a1a1b1a3b2   CTS8277
• • • • • • • • • • R1a1a1b1a3c   CTS4027
• • • • • • • R1a1a1b2   F992/S202/Z93
• • • • • • • • R1a1a1b2*   -
• • • • • • • • R1a1a1b2a   F3105/S340/Z94, L342.2/S278.2
• • • • • • • • • R1a1a1b2a*   -
• • • • • • • • • R1a1a1b2a1   L657/S347
• • • • • • • • • • R1a1a1b2a1*   -
• • • • • • • • • • R1a1a1b2a1a   Y7
• • • • • • • • • • R1a1a1b2a1b   Y6
• • • • • • • • • R1a1a1b2a2   Z2124
• • • • • • • • • • R1a1a1b2a2*   -
• • • • • • • • • • R1a1a1b2a2a   Z2123
• • • • • • • • • • R1a1a1b2a2b   S4576/Z2122
• • • • • • • • • • • R1a1a1b2a2b*   -
• • • • • • • • • • • R1a1a1b2a2b1   F1345
• • • • • • • • • • • • R1a1a1b2a2b1*   -
• • • • • • • • • • • • R1a1a1b2a2b1a   CTS6/S3403
• • • • • • • • • • • • R1a1a1b2a2b1b   F2935
• • • • • • • • • • • R1a1a1b2a2b2   Y57

• • R1b   M343/PF6242


• • • R1b*   -
• • • R1b1   P25_1, P25_2, P25_3, L278, M415/PF6251
• • • • R1b1*   -
• • • • R1b1a   P297/PF6398, L320
• • • • • R1b1a*   -
• • • • • R1b1a1   M73, M478
• • • • • R1b1a2   M269, CTS8728/L1063/PF6480/S13, L265/PF6431, M520/PF6410, PF6475/S17/YSC0000269, PF6485/S3, PF6399/S10
• • • • • • R1b1a2*   -
• • • • • • R1b1a2a   L23/PF6534/S141, L49.1/S349, L150.1/PF6274.1/S351.1
• • • • • • • R1b1a2a*   -
• • • • • • • R1b1a2a1   L51/M412/PF6536/S167
• • • • • • • • R1b1a2a1*   -
• • • • • • • • R1b1a2a1a   L151/PF6542, L11/PF6539/S127, L52/PF6541, P310/PF6546/S129, P311/PF6545/S128
• • • • • • • • • R1b1a2a1a*   -

• • • • • • • • • R1b1a2a1a1   M405/U106/S21


• • • • • • • • • • R1b1a2a1a1*   -
• • • • • • • • • • R1b1a2a1a1a   FGC3861
• • • • • • • • • • • R1b1a2a1a1a*   -
• • • • • • • • • • • R1b1a2a1a1a1   L217.1/S1849.1
• • • • • • • • • • • R1b1a2a1a1a2   FGC14877
• • • • • • • • • • R1b1a2a1a1b   Z19, S493/YSC0000053/Z18
• • • • • • • • • • • R1b1a2a1a1b*   -
• • • • • • • • • • • R1b1a2a1a1b1   Z14
• • • • • • • • • • • • R1b1a2a1a1b1*   -
• • • • • • • • • • • • R1b1a2a1a1b1a   S375/Z372
• • • • • • • • • • • • • R1b1a2a1a1b1a*   -
• • • • • • • • • • • • • R1b1a2a1a1b1a1   L257/S186
• • • • • • • • • • • • • • R1b1a2a1a1b1a1*   -
• • • • • • • • • • • • • • R1b1a2a1a1b1a1a   S5741, L729.2/M2087.2/Z15.2/Z548.2, Z375, Z378/S5757
• • • • • • • • • • • • R1b1a2a1a1b1b   DF95/S21484
• • • • • • • • • • R1b1a2a1a1c   S263/Z381
• • • • • • • • • • • R1b1a2a1a1c*   -
• • • • • • • • • • • R1b1a2a1a1c1   S264/Z156
• • • • • • • • • • • • R1b1a2a1a1c1*   -
• • • • • • • • • • • • R1b1a2a1a1c1a   S497/Z306, S265/Z304, S376/Z305, S498/Z307
• • • • • • • • • • • • • R1b1a2a1a1c1a*   -
• • • • • • • • • • • • • R1b1a2a1a1c1a1   DF98
• • • • • • • • • • • • • R1b1a2a1a1c1a2   DF96/S1809
• • • • • • • • • • • • • • R1b1a2a1a1c1a2*   -
• • • • • • • • • • • • • • R1b1a2a1a1c1a2a   L1/S26
• • • • • • • • • • • • • • R1b1a2a1a1c1a2b   P89.2
• • • • • • • • • • • • R1b1a2a1a1c1b   S5520
• • • • • • • • • • • R1b1a2a1a1c2   S499/Z301
• • • • • • • • • • • • R1b1a2a1a1c2*   -
• • • • • • • • • • • • R1b1a2a1a1c2a   M467/S29/U198
• • • • • • • • • • • • R1b1a2a1a1c2b   L48/S162
• • • • • • • • • • • • • R1b1a2a1a1c2b*   -
• • • • • • • • • • • • • R1b1a2a1a1c2b1   L47/S170
• • • • • • • • • • • • • • R1b1a2a1a1c2b1*   -
• • • • • • • • • • • • • • R1b1a2a1a1c2b1a   L44/S171, L163/S352
• • • • • • • • • • • • • • • R1b1a2a1a1c2b1a*   -
• • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1   L46/S172
• • • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1a   L525
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1a*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1a1   L45/S353, L164/S502, L237
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1a1*   -
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b1a1a1a   L477, L493
• • • • • • • • • • • • • • R1b1a2a1a1c2b1b   Z159, Z160, Z350
• • • • • • • • • • • • • R1b1a2a1a1c2b2   S268/Z9, S379/Z10, S504/Z28, Z348
• • • • • • • • • • • • • • R1b1a2a1a1c2b2*   -
• • • • • • • • • • • • • • R1b1a2a1a1c2b2a   S271/Z30
• • • • • • • • • • • • • • • R1b1a2a1a1c2b2a*   -
• • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1   Z2, S511
• • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a   S272/YSC0000049/Z7, S509/Z31
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1   YSC0000050/Z8, S384/YSC0000047/Z5, S515/Z21, Z22, Z24, Z25, S517/Z26, Z29, S516/Z351
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1*   -
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1a   S274/Z11
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1a*   -
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1a1   S385/Z12
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1a1*   -
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1a1a   L148/S173
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b   S275/YSC0000043/Z1
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b*   -
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b1   S514/Z344
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b1*   -
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b1a   S276/Z6
• • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b1a*   -
• • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b1a1   S18951
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2   S512/Z346
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2*   -
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2a   S387/Z343
• • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2a*   -
• • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2a1   S3933
• • • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2a1*   -
• • • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2a1a   CTS7080/S3334
• • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2b   DF101
• • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2b*   -
• • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2b1   DF102
• • • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2b1*   -
• • • • • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1b2b1a   FGC12993
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a1c   M365.4
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a2   CTS10893/S3595
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a2*   -
• • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a2a   FGC924
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a2a*   -
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a2a1   FGC918
• • • • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2a1a2a2   A294
• • • • • • • • • • • • • • R1b1a2a1a1c2b2b   S1743/Z331, Z334, Z347
• • • • • • • • • • • • • • • R1b1a2a1a1c2b2b*   -
• • • • • • • • • • • • • • • R1b1a2a1a1c2b2b1   S505/Z330
• • • • • • • • • • • • • • • • R1b1a2a1a1c2b2b1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a1c2b2b1a   Z326, S380/Z329, Z337
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2b1a*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2b1a1   CTS2509/S1734, S1741/Z319, Z325
• • • • • • • • • • • • • • • • • R1b1a2a1a1c2b2b1a2   S21728
• • • • • • • • • • • • • R1b1a2a1a1c2b3   L200
• • • • • • • • • • R1b1a2a1a1d   FGC396
• • • • • • • • • • • R1b1a2a1a1d*   -
• • • • • • • • • • • R1b1a2a1a1d1   L199.1/L443.1/L675.1/PF5371.1
• • • • • • • • • • R1b1a2a1a1e   S12025

• • • • • • • • • R1b1a2a1a2   P312/PF6547/S116


• • • • • • • • • • R1b1a2a1a2*   -
• • • • • • • • • • R1b1a2a1a2a   DF27/S250
• • • • • • • • • • • R1b1a2a1a2a*   -
• • • • • • • • • • • R1b1a2a1a2a1   Z195/S227, S355/Z196
• • • • • • • • • • • • R1b1a2a1a2a1*   -
• • • • • • • • • • • • R1b1a2a1a2a1a   S230/Z209, S356/Z220
• • • • • • • • • • • • • R1b1a2a1a2a1a*   -
• • • • • • • • • • • • • R1b1a2a1a2a1a1   S181/Z278, Z216
• • • • • • • • • • • • • • R1b1a2a1a2a1a1*   -
• • • • • • • • • • • • • • R1b1a2a1a2a1a1a   S348/Z214
• • • • • • • • • • • • • • • R1b1a2a1a2a1a1a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2a1a1a1   M153
• • • • • • • • • • • • R1b1a2a1a2a1b   S228/Z198, L176.2/S179.2
• • • • • • • • • • • • • R1b1a2a1a2a1b*   -
• • • • • • • • • • • • • R1b1a2a1a2a1b1   S460/Z292
• • • • • • • • • • • • • • R1b1a2a1a2a1b1*   -
• • • • • • • • • • • • • • R1b1a2a1a2a1b1a   Z262
• • • • • • • • • • • • • • • R1b1a2a1a2a1b1a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2a1b1a1   M167/SRY2627
• • • • • • • • • • • • • R1b1a2a1a2a1b2   L165/S68
• • • • • • • • • • • • • R1b1a2a1a2a1b3   CTS4188
• • • • • • • • • • • • R1b1a2a1a2a1c   DF17/S455
• • • • • • • • • • • R1b1a2a1a2a2   Z2552
• • • • • • • • • • • • R1b1a2a1a2a2*   -
• • • • • • • • • • • • R1b1a2a1a2a2a   L617
• • • • • • • • • • • R1b1a2a1a2a3   L881
• • • • • • • • • • R1b1a2a1a2b   PF6570/S28/U152
• • • • • • • • • • • R1b1a2a1a2b*   -
• • • • • • • • • • • R1b1a2a1a2b1   L2/S139
• • • • • • • • • • • • R1b1a2a1a2b1*   -
• • • • • • • • • • • • R1b1a2a1a2b1a   S255/Z367
• • • • • • • • • • • • • R1b1a2a1a2b1a*   -
• • • • • • • • • • • • • R1b1a2a1a2b1a1   L20/S144
• • • • • • • • • • • • • R1b1a2a1a2b1a2   S368/Z34
• • • • • • • • • • • • • • R1b1a2a1a2b1a2*   -
• • • • • • • • • • • • • • R1b1a2a1a2b1a2a   S487/Z35
• • • • • • • • • • • • • • • R1b1a2a1a2b1a2a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2b1a2a1   Z275
• • • • • • • • • • • • R1b1a2a1a2b1b   L196
• • • • • • • • • • • • R1b1a2a1a2b1c   Z49
• • • • • • • • • • • • • R1b1a2a1a2b1c*   -
• • • • • • • • • • • • • R1b1a2a1a2b1c1   S211/Z142
• • • • • • • • • • • • • • R1b1a2a1a2b1c1*   -
• • • • • • • • • • • • • • R1b1a2a1a2b1c1a   L562/S213
• • • • • • • • • • • R1b1a2a1a2b2   S206/Z36
• • • • • • • • • • • R1b1a2a1a2b3   PF6601/Z56
• • • • • • • • • • • • R1b1a2a1a2b3*   -
• • • • • • • • • • • • R1b1a2a1a2b3a   L4/S178
• • • • • • • • • • • • R1b1a2a1a2b3b   S47
• • • • • • • • • • • • R1b1a2a1a2b3c   PF6578/S371/Z145, PF6584/S483/Z146, Z144
• • • • • • • • • • R1b1a2a1a2c   L21/M529/S145, L459, S461/Z290
• • • • • • • • • • • R1b1a2a1a2c*   -
• • • • • • • • • • • R1b1a2a1a2c1   CTS241/DF13/S521
• • • • • • • • • • • • R1b1a2a1a2c1*   -
• • • • • • • • • • • • R1b1a2a1a2c1a   DF49/S474
• • • • • • • • • • • • • R1b1a2a1a2c1a*   -
• • • • • • • • • • • • • R1b1a2a1a2c1a1   S6154/Z2980
• • • • • • • • • • • • • • R1b1a2a1a2c1a1*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1a1a   S476/Z2976
• • • • • • • • • • • • • • • R1b1a2a1a2c1a1a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1a1a1   DF23/S193
• • • • • • • • • • • • • • • • R1b1a2a1a2c1a1a1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a2c1a1a1a   Z2961
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1a1a*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1a1a1   M222/Page84/USP9Y+3636
• • • • • • • • • • • • • • • • • • R1b1a2a1a2c1a1a1a1*   -
• • • • • • • • • • • • • • • • • • R1b1a2a1a2c1a1a1a1a   DF85/S675
• • • • • • • • • • • • R1b1a2a1a2c1b   DF1/L513/S215
• • • • • • • • • • • • • R1b1a2a1a2c1b*   -
• • • • • • • • • • • • • R1b1a2a1a2c1b1   P66_1, P66_2, P66_3
• • • • • • • • • • • • • R1b1a2a1a2c1b2   L193.1/S176.1
• • • • • • • • • • • • • R1b1a2a1a2c1b3   L706.2
• • • • • • • • • • • • • • R1b1a2a1a2c1b3*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1b3a   L705.2/S465.2
• • • • • • • • • • • • • R1b1a2a1a2c1b4   CTS3087
• • • • • • • • • • • • R1b1a2a1a2c1c   L96
• • • • • • • • • • • • R1b1a2a1a2c1d   L195/S354, L144.1/S175
• • • • • • • • • • • • R1b1a2a1a2c1e   S219/Z255
• • • • • • • • • • • • • R1b1a2a1a2c1e*   -
• • • • • • • • • • • • • R1b1a2a1a2c1e1   L159.2/S169.2
• • • • • • • • • • • • R1b1a2a1a2c1f   S218/Z253
• • • • • • • • • • • • • R1b1a2a1a2c1f*   -
• • • • • • • • • • • • • R1b1a2a1a2c1f1   L554
• • • • • • • • • • • • • R1b1a2a1a2c1f2   S686/Z2534
• • • • • • • • • • • • • • R1b1a2a1a2c1f2*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1f2a   L226/S168
• • • • • • • • • • • • • • R1b1a2a1a2c1f2b   L643
• • • • • • • • • • • • • • R1b1a2a1a2c1f2c   S893/Z2185
• • • • • • • • • • • • • • • R1b1a2a1a2c1f2c*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1f2c1   CTS1202.1/L1066.1
• • • • • • • • • • • • • • • • R1b1a2a1a2c1f2c1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a2c1f2c1a   CTS9881
• • • • • • • • • • • • • • R1b1a2a1a2c1f2d   DF73/S923
• • • • • • • • • • • • • R1b1a2a1a2c1f3   PF825.2
• • • • • • • • • • • • R1b1a2a1a2c1g   DF21/S192
• • • • • • • • • • • • • R1b1a2a1a2c1g*   -
• • • • • • • • • • • • • R1b1a2a1a2c1g1   FGC3213
• • • • • • • • • • • • • • R1b1a2a1a2c1g1*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1g1a   P314.2/S220.2
• • • • • • • • • • • • • • • R1b1a2a1a2c1g1a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1g1a1   L294.2/L362.2/S312.2
• • • • • • • • • • • • • • R1b1a2a1a2c1g1b   S3058
• • • • • • • • • • • • • • • R1b1a2a1a2c1g1b*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1g1b1   S423, S424, S426, S3025, S3026, S3031, S3057
• • • • • • • • • • • • • • • • R1b1a2a1a2c1g1b1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a2c1g1b1a   CTS2187/S190, CTS9333/S307, S301, S308, S309, S427, S3027, S3028, S3034   (See Notes regarding S3028)
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1g1b1a*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1g1b1a1   S425, S3033
• • • • • • • • • • • • • R1b1a2a1a2c1g2   FGC3903/S5201/Y2890
• • • • • • • • • • • • • • R1b1a2a1a2c1g2*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1g2a   S280/Z246
• • • • • • • • • • • • • • • R1b1a2a1a2c1g2a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1   DF25/S253
• • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a   DF5/S191, S281/Z248
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a1   L658
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a2   CTS3655/S3787
• • • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a2*   -
• • • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a2a   L627
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a3   L1403
• • • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a3*   -
• • • • • • • • • • • • • • • • • • R1b1a2a1a2c1g2a1a3a   L1402
• • • • • • • • • • • • • R1b1a2a1a2c1g3   S5488
• • • • • • • • • • • • • • R1b1a2a1a2c1g3*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1g3a   S7200
• • • • • • • • • • • • • • • R1b1a2a1a2c1g3a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1g3a1   L720/S299
• • • • • • • • • • • • • • R1b1a2a1a2c1g3b   Z16294
• • • • • • • • • • • • • • • R1b1a2a1a2c1g3b*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1g3b1   L130
• • • • • • • • • • • • • R1b1a2a1a2c1g4   S5456
• • • • • • • • • • • • • R1b1a2a1a2c1g5   S971/Z3017
• • • • • • • • • • • • R1b1a2a1a2c1h   L371/S300
• • • • • • • • • • • • R1b1a2a1a2c1i    CTS2501/S836, CTS6581/DF41/S524
• • • • • • • • • • • • • R1b1a2a1a2c1i*   -
• • • • • • • • • • • • • R1b1a2a1a2c1i1   S775
• • • • • • • • • • • • • • R1b1a2a1a2c1i1*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1i1a   L744/S388, L745/S463, L746/S310
• • • • • • • • • • • • • R1b1a2a1a2c1i2   BY114/FGC14692
• • • • • • • • • • • • • • R1b1a2a1a2c1i2*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1i2a   L563
• • • • • • • • • • • • R1b1a2a1a2c1j   S470/Z251
• • • • • • • • • • • • • R1b1a2a1a2c1j*   -
• • • • • • • • • • • • • R1b1a2a1a2c1j1   L555/S393
• • • • • • • • • • • • R1b1a2a1a2c1k   L1335/S530, S730, S737, S747, S5264, S5280, S5284, S5285
• • • • • • • • • • • • • R1b1a2a1a2c1k*   -
• • • • • • • • • • • • • R1b1a2a1a2c1k1   CTS11722/L1065/S749, CTS3057, CTS6838, CTS7030/S735, CTS10008, CTS10351, M10953/S738
• • • • • • • • • • • • • • R1b1a2a1a2c1k1*   -
• • • • • • • • • • • • • • R1b1a2a1a2c1k1a   S744, Z16325
• • • • • • • • • • • • • • • R1b1a2a1a2c1k1a*   -
• • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1   S764, S7370
• • • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1*   -
• • • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1a   L743
• • • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1b   FGC17596/Z17611, FGC17597/Z17613
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1b*   -
• • • • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1b1   FGC17595/Z17612
• • • • • • • • • • • • • • • • R1b1a2a1a2c1k1a1c   S756, Z16327
• • • • • • • • • • • • • • • R1b1a2a1a2c1k1a2   S691/Z13851
• • • • • • • • • • • • • • R1b1a2a1a2c1k1b   Z16329, Z16330, Z16330
• • • • • • • • • • • • • • R1b1a2a1a2c1k1c   FGC10125
• • • • • • • • • • • • R1b1a2a1a2c1l   CTS4466/S1136
• • • • • • • • • • • • • R1b1a2a1a2c1l*   -
• • • • • • • • • • • • • R1b1a2a1a2c1l1   L270.2
• • • • • • • • • • • • R1b1a2a1a2c1m   CTS2457.2
• • • • • • • • • • • • R1b1a2a1a2c1n   L679
• • • • • • • • • • • R1b1a2a1a2c2   CTS300/DF63/S522
• • • • • • • • • • • • R1b1a2a1a2c2*   -
• • • • • • • • • • • • R1b1a2a1a2c2a   CTS6919/S3689
• • • • • • • • • • R1b1a2a1a2d   L238/S182
• • • • • • • • • • R1b1a2a1a2e   DF19/S232
• • • • • • • • • • • R1b1a2a1a2e*   -
• • • • • • • • • • • R1b1a2a1a2e1   DF88/S4298
• • • • • • • • • • • • R1b1a2a1a2e1*   -
• • • • • • • • • • • • R1b1a2a1a2e1a   L644
• • • • • • • • • • • • R1b1a2a1a2e1b   L1199
• • • • • • • • • • • • R1b1a2a1a2e1c   L719
• • • • • • • • • • • R1b1a2a1a2e2   S233/Z302
• • • • • • • • • • R1b1a2a1a2f   DF99/S11987

• • • • • • • • R1b1a2a1b   PF7589/Z2118


• • • • • • • • • R1b1a2a1b*   -
• • • • • • • • • R1b1a2a1b1   CTS6889/S1161
• • • • • • • • • • R1b1a2a1b1*   -
• • • • • • • • • • R1b1a2a1b1a   CTS11824/PF7592
• • • • • • • R1b1a2a2   CTS1078/Z2103, Z2105
• • • • • • • • R1b1a2a2*   -
• • • • • • • • R1b1a2a2a   L584
• • • • • • • • • R1b1a2a2a*   -
• • • • • • • • • R1b1a2a2a1   L943
• • • • • • • • R1b1a2a2b   L277.1/S334.1
• • • • • • • • R1b1a2a2c   L150.2!/PF6274.2!
• • • • • • • • R1b1a2a2d   CTS7822/Z2110
• • • • • • • • • R1b1a2a2d*   -
• • • • • • • • • R1b1a2a2d1   CTS9219
• • • • R1b1b   M335
• • • • R1b1c   PF6279/V88
• • • • • R1b1c*   -
• • • • • R1b1c1   M18
• • • • • R1b1c2   V35
• • • • • R1b1c3   V69

R2   M479/PF6107


• • R2*   -
• • R2a   L266/PF6108, M124, P249, P267/PF6109
• • • R2a*   -
• • • R2a1   L295
• • • • R2a1*   -
• • • • R2a1a   L294.1/L362.1/S312.1
• • • • R2a1b   L723
• • • • • R2a1b*   -
• • • • • R2a1b1   L725
• • • • • • R2a1b1*   -
• • • • • • R2a1b1a   L724
• • • R2a2   L263
• • • R2a3   L1069

Link to Haplogroup R Private SNPs, Notes and References (Papers)

Y-DNA haplogroup R-M207 is believed to have arisen approximately 27,000 years ago in Asia. The two currently defined subclades are R1 and R2.
R1-M173 is estimated to have arisen during the height of the Last Glacial Maximum (LGM), about 18,500 years ago, most likely in southwestern Asia. The two most common descendant clades of R1 are R1a and R1b.
R1a-M420 is believed to have arisen on the Eurasian Steppe or the Indus Valley, and today is most frequently observed in eastern Europe and in western and central Asia. R1a1a1g-M458 is found at frequencies approaching or exceeding 30% in Eastern Europe.
R1b-M343 is believed to have arisen in southwest Asia and today its sublcades are bound in various distributions across Eurasia and Africa.
Paragroup R1b1* and R1b1c-V88 are found most frequently in SW Asia and Africa. The African examples are almost entirely within R1b1c and are associated with the spread of Chadic languages.
R1b1a-P297 is found throughout Eurasia. R1b1a1-M73 is observed most frequently in Asia, with low frequency of observation in Europe. R1b1a2-M269 is observed most frequently in Europe, especially western Europe, but with notable frequency in southwest Asia.
R1b1a2-M269 is estimated to have arisen approximately 4,000 to 8,000 years ago in southwest Asia and to have spread into Europe from there. The Atlantic Modal Haplotype, or AMH, is the most common STR haplotype in R1b1a2a1a-L11/PF6539/S127 and most European R1b1a2 belongs to R1b1a2a1a1-M405/S21/U106 or R1b1a2a1a2-P312/PF6547/S116.
R2-M479 is most often observed in Asia, especially on the Indian sub-continent and in central Asia.

Additional Resources: (Projects arranged in the order that they appear on the tree.)

ISOGG Wiki - What you need to know about Genetic Genealogy.
Y-Haplogroup R* DNA Project, Vincent Vizacherro.
R-Arabia Y-DNA Project.
R1* Haplogroup Project, Vincent Vizacherro.
R1a International Y-DNA Project.
R1a1a and Subclades Y-DNA Project, Lukasz Lapinski, Lawrence Mayka, Marius Wilkoszewski, A. J. Levin, Anne M. Berge, Michal Milewski.
R1a WTY Project, Cyndi Rutledge.
Ashkenazi-Levite DNA Project (R1a1).
Y-DNA Haplogroup R1b - FTDNA vs. ISOGG Haplotrees, Diana Gale Matthiesen.
Kerchner's R1b and Subclades YDNA Haplogroup Project, Tibor Fehér and Vincent Tilroe.
The Jewish R1b Project, Elise Friedman, Sean Silver, Itzhak Epstein, Gil Bardige.
R1b1(xP297) DNA Project, Vincent Vizacherro, Peter Hrechdakian.
R1b1b1 (R1b1a1) Haplogroup Project (M73), Vincent Vizacherro.
R1b1a2 (P312- U106-) DNA Project (aka ht35 Project), Vincent Vizacherro, Peter Hrechdakian.
R1b1b2* (R1b1a2*) Project.
YDNA Haplogroup R1b-U106/S21+ Research Group, Charles Moore.
R-U106xL48 WTY Project.
The U198 / S29 Project, John Sloan.
R-L48+ WTY Project, Charles Moore.
R1b-P89.2, Gary Blakely.
R-P312 and Subclades Project, Richard Stevens.
R-M153 The Basque Marker.
R1b - U152 & Subclades Research Project, Tibor Fehér, Steve Gilbert.
U152+/S28+ Data, David Faux.
R-L21 Plus Project, Richard Stevens,
R-DF21 and Subclades, David Reynolds.
R-L21 Walk Through the Y, Kirsten Saxe, Vincent Tilroe, Alex Williamson.
R1b DYS464X=15.3g Subclade, David Reynolds.
R-L21* 11-13 Combo: includes L193+, P314.2+, Adam Bradford.
R-L21 4466 South Irish, Kathleen Kerwin.
R1b-CTS4466 Plus, Elizabeth O'Donoghue/Ross
R-M222 Haplogroup Project (Niall of the 9 Hostages, Ui Neill Clan), David Wilson.
R-L159 Project.
The R-L226 Project - Irish Type III, Dennis O'Brien, Dennis Wright.
The Irish Type III Website (L226) (Dalcassian Clans), Dennis Wright.
The Y-Haplogroup R SRY2627+ and L176.2+ Project, Gareth Henson.
R-L165 (S68) Project, Lori McLeod-Wilke, Alasdair Macdonald, Conrad Terrill, Timothy McLeod.
R-17-14-10 Haplogroup Project (aka L371), Robert Hughes.
R2 Y-Chromosome Haplogroup DNA Project.
R2-M124-WTY Project.

Corrections/Additions made since 1 January 2014:

  • Added S334.1, S340, S351.1, S3334, S3361, S3403, S3595 to tree; added S357, S358 to R1b Investigation on 6 January 2014.

  • Added PF6057, YSC0000201 to tree on 18 January 2014.

  • Added PF6410 on 19 January 2014.

  • Moved L745/S463 to a different subclade on tree; corrected errors to correctly read S275/YSC0000043/Z1, S272/YSC0000049/Z7 and S514/Z344; updated paragraph containing the words "Atlantic Modal Haplotype" on 22 January 2014.

  • Added CTS8728, L1063, PF6014, PF6475, PF6480, YSC0000269 on 23 January 2014.

  • Removed David Reynolds as contact person for R-P312 and Subclades on 30 January 2014.

  • Added PF3499, PF6050, PF6112, PF6223, PF6236, PF6240 to tree; added F820 to R2 Investigation on 30 January 2014.

  • Added S371, S379, S384/YC000047, S3689, S3787, S4298 to tree; moved CTS6838, CTS7030/S735 from R1b Investigation to tree; added S360, S362, S372, S691/Z13851, S3666, S4247 to Investigation on 31 January 2014.

  • Added CTS300, CTS6581, S461, S465.2, S470, S487, S497, S498, S499, S511, S4554, S4576, YSC0000050, YSC0000053 to tree; added S453, S482. S637, S640, S641, S651, S653 to R1b Investigation on 1 February 2014.

  • Added PF6234, S836, S868 to tree; added S883, S893, S8070 to R1b Investigation on 2 February 2014.

  • Added PF7598 to Investigation on 3 February 2014.

  • Added PF6636 to Investigation on 4 February 2014.

  • Removed link to Dal Riadic Migration Y Chromosome DNA Genealogy Page (L21), John McEwan as it is no longer active on 6 February 2014.

  • Added R1b-CTS4466 Plus to Additional Resources on 6 February 2014.

  • Added L270.2 to tree; added CTS8358 to R1b Investigation on 6 February 2014.

  • Changed R1b-P312 contact person to Mark Jost on 6 February 2014.

  • Added CTS9881 to tree; added R-L21 4466 South Irish to Additional Resources on 13 February 2014.

  • Added CTS9333/S307, S301, S423, S425, S3025, S3026, S3027, S3028, S3031, S3033, S3034, S3057, S3058 to tree; added S3106 to R1b Investigation; added S3068 to R1b Private; moved S303, S421 from R1b Investigation to R1b Private on 23 February 2014.

  • Changed subclade names from S3058 and downstream on 25 February 2014.

  • Added F3105 on 5 March 2014.

  • Added FGC5496, M1313.2, S588/FGC4108, S658/DF106, S659/DF105, S660/DF109, S661/DF104, S7073/FGC462, S7814 to R1b Investigation on 7 March 2014.

  • Added Z2355 to R1b Investigation on 13 March 2014.

  • Moved DF95/S21484 from R1b Investigation to tree on 16 March 2014.

  • Removed CTS358.2, CTS7275, CTS8127.2, YSC0000081.2 from R1b Investigation on 28 March 2014.

  • Moved Z334 from R1b Investigation to tree; added Z347 to tree on 29 March 2014.

  • Added Y6 to tree on 30 March 2014.

  • Moved S380/Z329 from R1b Investigation to tree; added DF101, DF102, S265/Z304, S504/Z28, S509/Z31, Z21, Z22, Z24, Z25, Z26, Z29, Z325, Z337, Z348, Z351, Z160, Z350 to tree on 31 March 2014.

  • Added S516, S517 to tree on 4 April 2014.

  • Added M734 on 26 April 2014.

  • Added A294, FGC396, FGC918, FGC924, FGC3861, FGC12993, S3933, S5520, S18951, S21728 to tree on 12 May 2014.

  • Added L525 to tree; moved L128/PF3499 off the tree to Frequently Mutating SNPs on 13 May 2014.

  • Moved L147 to Frequently Mutating SNPs on 14 May 2014.

  • Added S12025 to tree on 21 May 2014.

  • Corrected error in R1; P294 is identical to PF6112 (not PF6014). Corrected on 16 May 2014.

  • Added M3268.2 to R1b Investigation on 14 August 2014.

  • Added CTS6889/S1161, CTS7822/Z2110, FGC14877, PF7589/Z2118 to tree on 27 August 2014.

  • Added Z2572 to R1b Investigation on 12 September 2014.

  • Added Underhill et al (2014) on 12 September 2014.

  • Added CTS11824/PF7592 to tree on 14 September 2014.

  • Added CTS9219 to tree on 18 September 2014.

  • Added .2 to Z15 in R1b Investigation; added L729.2/M2087.2/Z15.2/Z548.2 to R1b Investigation on 18 September 2014.

  • Moved L1403 from R1b Investigation to tree on 20 September 2014.

  • Added .1 and .2 to L297 and L362 and added S312.1, S312.1; added L443.1/L675.1/PF5371.1, Z2552 to tree; added L199.3/L675.3/PF5371.3 to Investigation on 30 September 2014.

  • Added S5741, Z375, Z378/S5757, ZP1 to tree; moved L729.2/M2087.2/Z15.2/Z548.2 from R1b Investigation to tree on 1 October 2014.

  • Added Charles Fueston as a Contact person for R1b-P312->DF21 on 1 October 2014.

  • Added CTS3057, CTS10008, CTS10351, FGC17595/Z17612, FGC17596/ Z17611, FGC17597/Z17613, M10953/S738, S730, S737, S744, S747, S756, S764, S5264, S5280, S5284, S5285, S7370, Z16325 to tree on 2 October 2014.

  • Added FGC3213, FGC3903/S5201/Y2890, FGC10125, S5488, S7200, Z16327, Z16329, Z16330, Z16330 to tree; moved S691/Z13851 from R1b Investigation to tree on 3 October 2014.

  • Added Stephen Parrish as a contact person; added BY114/FGC14692, S775, S971/Z3017, S5456, Z16294 to tree on 4 October 2014.

  • Added S476/Z2976, S6154/Z2980 to tree on 6 October 2014.

  • Added David Stedman as a contact person on 7 October 2014.

  • Moved S228/Z198 from Investigation to tree on 8 October 2014.

  • Added Kathleen Kerwin as a contact person on 8 October 2014.

  • Added S460/Z292 to tree on 17 October 2014.

  • Added L943 to tree on 18 October 2014.

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CTS10168+, CTS10362+, CTS10834+, CTS109+, CTS11358+, CTS11468+, CTS11575+, CTS11726+, CTS11985+, CTS12478+, CTS125+, CTS12632+, CTS1996+, CTS2134+, CTS2664+, CTS3063+, CTS3135+, CTS3331+, CTS3358+, CTS3431+, CTS3536+, CTS3575+, CTS3654+, CTS3662+, CTS3868+, CTS3996+, CTS4244+, CTS4364+, CTS4368+, CTS4437+, CTS4443+, CTS4740+, CTS5318+, CTS5457+, CTS5532+, CTS5577+, CTS5884+, CTS6135+, CTS623+, CTS6383+, CTS6800+, CTS6907+, CTS7400+, CTS7659+, CTS7922+, CTS7933+, CTS8243+, CTS8591+, CTS8665+, CTS8728+, CTS8980+, CTS9828+, F1046+, F115+, F1209+, F1302+, F1320+, F1329+, F1704+, F1714+, F1753+, F1767+, F1794+, F180+, F2048+, F2075+, F211+, F2142+, F2155+, F2302+, F2402+, F2587+, F2688+, F2710+, F2837+, F29+, F295+, F2985+, F2993+, F3111+, F313+, F3136+, F33+, F332+, F3335+, F344+, F3556+, F356+, F359+, F3692+, F378+, F4+, F47+, F506+, F556+, F63+, F640+, F647+, F652+, F671+, F719+, F82+, F83+, F93+, L11+, L132+, L15+, L150+, L151+, L16+, L23+, L265+, L278+, L350+, L388+, L389+, L407+, L468+, L470+, L471+, L478+, L482+, L483+, L498+, L500+, L502+, L506+, L51+, L52+, L585+, L721+, L747+, L752+, L754+, L761+, L768+, L773+, L774+, L779+, L82+, M139+, M168+, M207+, M235+, M294+, M343+, M415+, M42+, M45+, M526+, M89+, M94+, P128+, P131+, P132+, P135+, P136+, P138+, P14+, P141+, P145+, P146+, P148+, P151+, P158+, P159+, P160+, P166+, P187+, P207+, P225+, P226+, P228+, P229+, P230+, P232+, P233+, P235+, P236+, P237+, P238+, P240+, P242+, P243+, P244+, P245+, P280+, P281+, P282+, P283+, P284+, P285+, P286+, P295+, P297+, P310+, PAGES00083+, PF1016+, PF1029+, PF1031+, PF1040+, PF1046+, PF1061+, PF1092+, PF1097+, PF110+, PF1203+, PF1269+, PF1276+, PF192+, PF210+, PF212+, PF223+, PF234+, PF258+, PF2591+, PF2593+, PF2599+, PF2608+, PF2611+, PF2615+, PF2624+, PF263+, PF2643+, PF272+, PF2745+, PF2747+, PF2748+, PF2749+, PF2770+, PF278+, PF292+, PF316+, PF325+, PF342+, PF500+, PF5465+, PF5466+, PF5468+, PF5471+, PF5851+, PF5853+, PF5854+, PF5865+, PF5869+, PF5871+, PF5882+, PF5886+, PF5887+, PF5888+, PF5953+, PF5956+, PF5957+, PF5964+, PF5965+, PF5982+, PF6063+, PF6091+, PF6145+, PF6246+, PF6249+, PF6250+, PF6263+, PF6265+, PF6270+, PF6271+, PF6272+, PF6404+, PF6409+, PF6411+, PF6424+, PF6425+, PF6430+, PF6432+, PF6434+, PF6438+, PF6443+, PF6463+, PF6494+, PF6495+, PF6498+, PF6500+, PF6506+, PF6507+, PF6509+, PF6524+, PF667+, PF719+, PF725+, PF779+, PF796+, PF803+, PF815+, PF821+, PF840+, PF844+, PF892+, PF937+, PF951+, PF954+, PF970+, s10+, s3+, U106+, V186+, V189+, V205+, V52+, V9+, YSC0000067+, YSC0000072+, YSC0000075+, YSC0000082+, YSC0000166+, YSC0000176+, YSC0000179+, YSC0000182+, YSC0000186+, YSC0000191+, YSC0000194+, YSC0000201+, YSC0000203+, YSC0000205+, YSC0000207+, YSC0000213+, YSC0000219+, YSC0000224+, YSC0000225+, YSC0000227+, YSC0000230+, YSC0000232+, YSC0000233+, YSC0000251+, YSC0000269+, YSC0000270+, YSC0000279+, YSC0000288+, YSC0000294+, Z1+, Z2+, Z20+, Z22+, Z23+, Z24+, Z25+, Z26+, Z28+, Z29+, Z30+, Z31+, Z344+, Z348+, Z352+, Z381+, Z4+, Z6+, Z7+, Z8+, Z9+

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