National Forests in Florida Final Report


I.A.8.N.g. Saturated temperate or subpolar needle-leaved evergreen forest



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I.A.8.N.g. Saturated temperate or subpolar needle-leaved evergreen forest


I.A. Evergreen forest

I.A.8.N.g.2 Atlantic White-cedar Saturated Forest Alliance (A.196)


CHAMAECYPARIS THYOIDES SATURATED FOREST ALLIANCE

Alliance Concept

Summary: These Chamaecyparis thyoides-dominated forests are found on saturated, peaty substrates. Chamaecyparis thyoides tolerates only brief inundation. The canopy of forests in this alliance may be heavily dominated by Chamaecyparis thyoides, or they may contain pines (Pinus elliottii var. elliottii in Florida basins, Pinus serotina in North Carolina peat domes) or hardwoods (for instance Acer rubrum, Nyssa biflora, and Persea palustris, or Liriodendron tulipifera in one rare example). The subcanopy may contain Acer rubrum var. trilobum, Nyssa biflora, Magnolia virginiana, Nyssa biflora, Persea palustris, and Cliftonia monophylla. Sabal palmetto and Serenoa repens are abundant on hummocks in one Florida association. The shrub stratum ranges from open to dense and may contain Clethra alnifolia, Cyrilla racemiflora, Gaylussacia frondosa (= var. frondosa), Ilex coriacea, Ilex glabra, Lyonia lucida, Magnolia virginiana, Persea palustris, Smilax laurifolia, and Vaccinium formosum. In some examples, the herbaceous stratum is well-developed and may contain Orontium aquaticum, Mitchella repens, Drosera sp., Osmunda cinnamomea, and Osmunda regalis var. spectabilis; in others it tends to be sparse and includes Woodwardia areolata, Woodwardia virginica, and Osmunda cinnamomea. Sphagnum spp., other mosses, and lichens may be common on exposed peat and rotting wood. ^Examples of this alliance may be found along small blackwater or spring-fed streams not subject to much flooding or siltation (in Florida and the East Gulf Coastal Plain); along streams or at seepages (in the Fall-line Sandhills); or in Panhandle Florida in large, constantly saturated basins (inundated under several feet of water during the spring) where fire is an infrequent event; as well as peat dome forests of North Carolina and Virginia dominated by Chamaecyparis and Pinus serotina. Soils may be acidic peats but are usually circumneutral sands or sands overlain by peat. Soils are generally organic Histosols, composed of sand and peat, are permanently saturated, and include Histosols (Saprists and Hemists); the pH may be circumneutral or slightly acidic. Fires are infrequent and less destructive than in the northern range of Chamaecyparis thyoides. ^Gap succession is more important in this community than post-fire regeneration and accounts for the mixed, uneven-aged stands. Lightning strikes are the main cause of mortality in mature white-cedar trees within the range of this association. Atlantic white-cedar communities are early successional but Chamaecyparis thyoides is a long-lived species (250+ years); gap regeneration is hypothesized to be the primary means of reproduction. The community often succeeds to itself following fire or other disturbance. In the absence of fire and adequate gap regeneration, Atlantic White-cedar Forests may be replaced by Bay Forest. Known examples are generally neither flood-prone, nor exposed to frequent fire. One association is thought to become established following fire, with a return time of 25-100+ years.

Synonymy:

  • IIA2b. Atlantic White Cedar Swamp Forest, in part (Allard 1990)

  • Atlantic White Cedar Swamp Forest (Oberholster 1993)

  • Bottomland Forest, in part (FNAI 1992a)

  • White Cedar Swamp Forest (Wieland 1994b)

  • Peatland Atlantic White Cedar Forest (Schafale and Weakley 1990)

  • Streamhead Atlantic White Cedar Forest (Schafale and Weakley 1990)

  • Atlantic White Cedar Swamp (Nelson 1986)

  • Mesotrophic Saturated Forest, in part (Rawinski 1992)

  • Atlantic White-Cedar: 97, in part (Eyre 1980)

Comments: Clewell and Ward (1987) describe a community along backswamps of larger rivers in Florida and along the Gulf Coast where "flooding is nominal." This could represent an additional association.

Alliance Distribution

Range: This alliance is found in Alabama, Florida, Georgia, Mississippi, North Carolina, South Carolina, and Virginia, and possibly Delaware (?) and Maryland (?).

States: AL CT DE? FL GA MA ME MS NC NH NJ NY RI SC VA

USFS Ecoregions: 212D:P, 221A:C, 221B:C, 221D:C, 232A:C, 232B:C, 232C:C, 232D:C, M212B:?

Federal Lands: DOD (Dare County Bombing Range, Eglin, Fort Bragg, Fort Gordon); USFS (Apalachicola, Conecuh, De Soto, Ocala); USFWS (Alligator River, Great Dismal Swamp, Mississippi Sandhill Crane, Pocosin Lakes, St. Marks)

Alliance Sources

References: Allard 1990, Clewell 1971, Clewell and Ward 1987, Eyre 1980, FNAI 1992a, FNAI 1992b, Fleming 1998, Frost 1987, Frost and Musselman 1987, Korstian and Brush 1931, Laderman 1989, Landaal 1978, Moore and Carter 1987, Nelson 1986, Oberholster 1993, Rawinski 1992, Schafale and Weakley 1990, Terwilliger 1987, Wharton 1978, Wharton et al. 1982, Whitehead 1972, Wieland 1994b, Wiseman 1986

I.B.2.N.a. Lowland or submontane cold-deciduous forest


I.B. Deciduous forest

I.B.2.N.a.16 American Beech - White Oak Forest Alliance (A.228)


FAGUS GRANDIFOLIA - QUERCUS ALBA FOREST ALLIANCE

Alliance Concept

Summary: Dry-mesic to mesic forests with admixtures of Quercus alba and Fagus grandifolia that typically occur on mesic slopes and small stream bottoms in the Coastal Plain, and also in other adjacent physiographic provinces, including the southern part of Crowley's Ridge, Arkansas. This alliance is distributed primarily north of the distribution of Magnolia grandiflora. The canopy may include Liriodendron tulipifera, Liquidambar styraciflua, Quercus michauxii, Quercus pagoda, Carya cordiformis, Fraxinus americana, and Ulmus spp. In the southern part of the range, examples of this alliance may have Acer barbatum and Acer leucoderme in the subcanopy; other associates may include Acer rubrum, Carya alba, Carya myristiciformis, Carya ovata, Carya texana, Celtis laevigata, Diospyros virginiana, Fraxinus americana, Gleditsia triacanthos, Gymnocladus dioicus, Juglans cinerea, Liquidambar styraciflua, Morus rubra, Nyssa sylvatica, Prunus serotina var. serotina, Quercus michauxii, Quercus stellata, Ulmus americana, Ulmus alata, and Ulmus rubra. Magnolia grandiflora and Magnolia acuminata may occur on moister, lower slopes, or in the subcanopy. The total canopy cover is usually dense. Phoradendron leucarpum, Tillandsia usneoides, and Pleopeltis polypodioides ssp. michauxiana (= Polypodium polypodioides var. michauxianum) may occur as epiphytes on the canopy trees. The understory and tall-shrub strata may include Aesculus pavia var. pavia, Aralia spinosa, Asimina triloba, Ostrya virginiana, Carpinus caroliniana ssp. caroliniana, Cornus florida, Cercis canadensis, Ilex opaca var. opaca, Styrax grandifolius, Crataegus spathulata, and Crataegus marshallii. The short-shrub stratum, which may vary widely in diversity, may contain Amelanchier arborea, Callicarpa americana, Chionanthus virginicus, Crataegus marshallii, Frangula caroliniana (= Rhamnus caroliniana), Hydrangea arborescens, Hydrangea quercifolia, Ilex ambigua, Ilex opaca var. opaca, Lindera benzoin, Vaccinium virgatum, Vaccinium arboreum, Vaccinium elliottii, Viburnum acerifolium, Viburnum dentatum, and Viburnum rufidulum; it may be patchy. The herbaceous layer, which is typically sparse, may contain Athyrium filix-femina ssp. asplenioides, Botrychium spp., Diplazium pycnocarpon, Phegopteris hexagonoptera, Osmunda spp., Polystichum acrostichoides, Actaea pachypoda, Arisaema triphyllum, Symphyotrichum drummondii (= Aster drummondii), Chasmanthium sessiliflorum, Cynoglossum virginianum, Desmodium nudiflorum, Galium circaezans, Helianthus hirsutus, Lilium michauxii, Lithospermum tuberosum, Maianthemum racemosum, Mitchella repens, Pedicularis canadensis, Podophyllum peltatum, Polygonatum biflorum, Scleria oligantha, Smilax herbacea, Smilax pumila, Solidago auriculata, Spigelia marilandica, Tipularia discolor, Tragia cordata, Uvularia perfoliata, Vicia minutiflora, and Viola walteri. This alliance typically occurs on mesic calcareous silty clays, silty loams and silty clay loams, as well as loamy sands or loamy fine sands. This broad-leaved forest is known predominantly from mesic middle and lower slopes; in southeastern Texas it occupies ravines and ridges within creek bottoms. Associated geology includes the Cook Mountain and Jackson formations. In the northwestern part of the range, this association is found on deep loessal soils of Crowley's Ridge, Arkansas (Cross County south through Phillips County), canopies are dominated by Fagus grandifolia, Quercus alba, and Liriodendron tulipifera, with associates including Fraxinus americana, Sassafras albidum, Ulmus rubra, Quercus michauxii, Acer saccharum, Magnolia acuminata, Carya illinoinensis. and Liquidambar styraciflua. This alliance concept includes, at least tentatively, forests in Virginia which are dominated by Fagus grandifolia, Quercus muehlenbergii, and Acer barbatum. Some less diverse examples in the northeastern part of the range, in North Carolina, contain a canopy of Fagus grandifolia and Liquidambar styraciflua over a subcanopy with Oxydendrum arboreum and a (possibly dense) shrub layer of Kalmia latifolia. An additional less diverse example in South Carolina contains Fagus grandifolia, Quercus nigra, and Liquidambar styraciflua as canopy dominants over Ilex opaca and a well-developed shrub layer with Rhododendron canescens, Euonymus americana, Vaccinium elliottii, Symplocos tinctoria, Arundinaria gigantea, Asimina triloba, Callicarpa americana, and others.

Synonymy:

  • IA8d. Southern Mixed Hardwood Forest, in part (Allard 1990)

  • IA8b. Coastal Plain Calcareous Forest, in part (Allard 1990)

  • Coastal Plain Beech Forest (Foti 1994b)

  • Mixed Mesophytic Forest, in part (Foti 1994b)

  • Piedmont Mesic Broad-leaved Deciduous Forest (Ambrose 1990a)

  • Deep soil mesophytic forest, in part (Evans 1991)

  • Calcareous mesophytic forest, in part (Evans 1991)

  • Coastal Plain mesophytic cane forest, in part (Evans 1991)

  • Calcareous Forest, in part (Smith 1996a)

  • Hardwood Slope Forest, in part (Smith 1996a)

  • Basic Mesic Forest, Piedmont Subtype (Schafale and Weakley 1990)

  • Basic Mesic Forest, Coastal Plain Subtype (Schafale and Weakley 1990)

  • Piedmont/Coastal Plain Heath Bluff, in part (Schafale and Weakley 1990)

  • Fagus grandifolia forest alliance (Hoagland 1998a)

  • T1B2aI1a. Fagus grandifolia - Ilex opaca (Foti et al. 1994)

  • Beech-Sweet Gum-Tulip Poplar MAP (Pyne 1994)

  • American Beech-White Oak Series (Diamond 1993)

  • American Beech-Southern Magnolia Series, in part (Diamond 1993)

  • American Beech - White Oak / Mitchella Loamy Moist-Mesic Steep Slopes and Ravines, in part (Turner et al. 1999)

Comments:

Alliance Distribution

Range: This alliance is found in Alabama, Arkansas, Florida (?), Georgia, Kentucky, Louisiana, Mississippi, North Carolina, Oklahoma, South Carolina, Tennessee, Texas, and Virginia.

States: AL AR FL? GA KY LA MS NC OK SC TN TX VA

USFS Ecoregions: 222A:?, 222C:C, 222D:?, 231A:C, 231B:C, 231C:C, 231E:C, 232A:C, 232B:C, 232C:C, 232E:C, 232F:C, 234A:C

Federal Lands: COE (Claiborne Lake); DOD (Fort Benning, Fort Stewart); NPS (Colonial, Congaree Swamp, Kings Mountain); USFS (Angelina, Apalachicola?, Bienville, Conecuh, Croatan, Davy Crockett?, De Soto, Delta?, Francis Marion, Homochitto, Kisatchie, Oconee, Ouachita?, Sabine, St. Francis, Sam Houston?, Talladega, Tuskegee, Tombigbee); USFWS (Lake Isom)

Alliance Sources

References: Allard 1990, Ambrose 1990a, Clark 1974, Clark 1977c, Diamond 1993, Evans 1991, Foti 1994b, Foti et al. 1994, Frost et al. 1990, Hill 1992, Hoagland 1998a, Martin and Smith 1991, Monk 1965, Monk et al. 1989, Pyne 1994, Quarterman and Keever 1962, Rice and Peet 1997, Schafale and Weakley 1990, Smith 1996a, Soblo 1989, Turner et al. 1999, Ware 1970, Ware 1988, Ware and Ware 1992


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