Swainson, 1830 subtribe iolaina riley, 1958 Iolaus pallene. Photo courtesy Jeremy Dobson


Subgenus Philiolaus Stempffer & Bennett, 1958



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Subgenus Philiolaus Stempffer & Bennett, 1958



Bulletin de l’Institut Français d’Afrique Noire (A) 20: 1336 (1243-1347). Type-species: Iolaus parasilanus Rebel, by original designation. Synonymised with Argiolaus by Heath, 1983 (Arnoldia. Zimbabwe 9: 145-164) but used by Collins et al., (2003) for three new species described by them.

Iolaus (Philiolaus) parasilanus Rebel, 1914
Iolaus parasilanus Rebel, 1914. Annalen des (K.K.) Naturhistorischen Museums. Wien 28: 266 (219-294).

Type locality: Democratic Republic of Congo: “Aus dem Urwalde am Nordwestufer des Tanganika-Sees”.

Distribution: Ghana, Nigeria, Gabon, Congo, Angola, Democratic Republic of Congo, Uganda, Kenya, Tanzania.

Common name: Friendly sapphire.

Habitat: Forest.

Habits: Tends to stay high up in the forest canopy but is occasionally attracted to flowers lower down (Larsen, 1991).

Early stages: Nothing published.

Larval food:

Loranthus sp. (Loranthaceae) [Van Someren, 1974: 328].

Phragmanthera usuiensis usuiensis (Oliv.) M.G. Gilbert (Loranthaceae) [Congdon and Collins, 1998: 96].

Phragmanthera polycrypta (F. Didr.) Balle (Loranthaceae) [Congdon and Collins, 1998: 96].

Phragmanthera brieyi (De Wild) Polh. & Wiens (Loranthaceae) [Congdon and Bampton, 2000: 35].
Iolaus (Philiolaus) parasilanus parasilanus Rebel, 1914
Iolaus parasilanus Rebel, 1914. Annalen des (K.K.) Naturhistorischen Museums. Wien 28: 266 (219-294).

Type locality: Democratic Republic of Congo: “Aus dem Urwalde am Nordwestufer des Tanganika-Sees”.

Distribution: Democratic Republic of Congo (South Kivu), Tanzania (west).
Iolaus (Philiolaus) parasilanus divaricatus (Riley, 1928)
Argiolaus parasilanus divaricatus Riley, 1928. Novitates Zoologicae 34: 377 (374-394).

Type locality: Kenya: “Kenya Colony: Nandi Plateau, 5,700-6,200 feet”.

Distribution: Democratic Republic of Congo (Uele, Ituri, North Kivu, Tshopo), Uganda, Kenya (west), Tanzania (north-west).

Specific localities:

Kenya – Soy; Kakamega; Yala River; Nandi (Larsen, 1991).

Tanzania – Kere Hill, Minziro Forest (rare); Munene Forest (single female); Kikuru Forest (Congdon and Collins, 1998).
Iolaus (Philiolaus) parasilanus mabillei (Riley, 1928)
Argiolaus parasilanus mabillei Riley, 1928. Novitates Zoologicae 34: 378 (374-394).

Type locality: Angola: “Portuguese Congo, Landana”.

Distribution: Gabon, Congo, Angola (Cabinda), Democratic Republic of Congo (Equateur).
Iolaus (Philiolaus) parasilanus maesseni (Stempffer & Bennett, 1958)
Iolaphilus (?) parasilanus maesseni Stempffer & Bennett, 1958. Bulletin de l’Institut Français d’Afrique Noire (A) 20: 1340 (1243-1347).

Type locality: Ghana: “Togo britannique: Ho”.

Distribution: Ghana, Nigeria (south and Cross River loop).

Specific localities:

Ghana – Ho (TL); Akosombo Dam (Larsen, 2005a)

Nigeria – Agege (Larsen, 2005a); Omo (Larsen, 2005a); Benin (Larsen, 2005a); Okwangwo (Larsen, 2005a); Oban Hills (Larsen, 2005a); Omo Forest (Larsen, 2005a).

Iolaus (Philiolaus) ismenias (Klug, 1834)
Lycaena ismenias Klug, 1834 in Klug, 1829-45. In: Ehrenburg, C.G., Symbolae Physicae, seu icones et descriptiones corporum naturalium novarum aut minus cognitorum: pl. 40 ([183] pp.).

Type locality: Sudan: “Ambukohl”.

Distribution: Senegal, Guinea-Bissau, Burkina Faso, Ivory Coast (Warren-Gash, pers. comm., 2002), Ghana, Togo, Nigeria, Cameroon, Democratic Republic of Congo, Sudan, Uganda, Ethiopia.

Common name: White sapphire.

Habitat: Dry savanna (Collins et al., 2003). In West Africa it occurs mainly in Sudan savanna but also occurs in the garden suburbs of some cities (Larsen, 2005a).

Early stages: The eggs are laid singly (Larsen, 2005a).

Larval food:

Loranthus species (sensu lato) (Loranthaceae) [Larsen, 2005a].
Iolaus (Philiolaus) ismenias ismenias (Klug, 1834)
Lycaena ismenias Klug, 1834 in Klug, 1829-45. In: Ehrenburg, C.G., Symbolae Physicae, seu icones et descriptiones corporum naturalium novarum aut minus cognitorum: pl. 40 ([183] pp.).

Type locality: Sudan: “Ambukohl”.

Distribution: Senegal, Guinea-Bissau, Burkina Faso, Ivory Coast (Warren-Gash, pers. comm., 2002), Ghana, Togo, Nigeria (north), Cameroon (north), Democratic Republic of Congo, Sudan (south), Uganda.

Specific localities:

Ivory Coast – Abidjan (Larsen, 2005a).

Ghana – Accra (Larsen, 2005a).
Iolaus (Philiolaus) ismenias piaggiae Oberthür, 1883
Iolaus ismenias piaggiae Oberthür, 1883. Annali del Museo Civico di Storia Naturale (di Genova) Giacomo Doria 18: 730 (709-740).

Type locality: Ethiopia: “Kolla di Giagaguè-Agher”.

Distribution: Ethiopia.

Note: Larsen (2005a: 195) is certain that this taxon is specifically distinct but does not formally emend its status.

Iolaus (Philiolaus) christofferi Collins & Larsen, 2003
Iolaus christofferi Collins & Larsen, 2003. Metamorphosis 14 (3): 79 (63-110).

Type locality: Cameroon: “Wak, Ngaoundere, northern Cameroun, i.2000”. Holotype (male) in African Butterfly Research Institute, Nairobi; genitalia SCC 458.

General remarks: Named for the great Swedish lepidopterist of the 19th and early 20th Centuries, P.O. Christoffer Aurivillius (Collins et al., 2003).

Distribution: Cameroon (north) (known only from the type locality).

Habitat: Dry savanna (Collins et al., 2003).

Early stages: Nothing published.

Larval food: Nothing published.

Iolaus (Philiolaus) vansomereni (Stempffer & Bennett, 1958)
Iolaphilus vansomereni Stempffer & Bennett, 1958. Bulletin de l’Institut Français d’Afrique Noire (A) 20: 1319 (1243-1347).

Type locality: Uganda: “Metu Hills, Nil blanc, N.W. Madi, Ouganda”.

Distribution: Uganda (north-west).

Habitat: Savanna (Collins et al., 2003).

Early stages: Nothing published.

Larval food: Nothing published.

Iolaus (Philiolaus) newporti Larsen, 1994
Iolaus (Iolaphilus) newporti Larsen, 1994. Entomologist's Record and Journal of Variation 106 (11-12): 197 (197-200).



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