Fagus grandifolia - Quercus alba - Liriodendron tulipifera / Hydrangea arborescens / Schisandra glabra Forest (KP

American Beech - White Oak - Tuliptree / Smooth Hydrangea / Magnolia-vine Forest

American Beech - White Oak / American Holly / Southern Lady Fern Forest

American Beech - Red Oak - White Oak Forest Alliance

CONCEPT: Forests in this alliance occur in non-montane or low mountain mesic situations and are dominated by Fagus

grandifolia with or without some combination of the Quercus spp. Associated canopy and subcanopy species can include

Liriodendron tulipifera, Acer saccharum, Magnolia tripetala, Magnolia acuminata (Ozarks), Tilia americana (Ozarks), Quercus

muehlenbergii, Acer rubrum, Cornus florida, Ostrya virginiana, Aesculus sylvatica, and Ilex opaca. Some of these forests,

particularly in the Piedmont of South Carolina or in Arkansas, may have Acer barbatum. Shrubs in this alliance include

herb layer can be relatively lush with such species as Polystichum acrostichoides, Galium circaezans, Hexastylis arifolia,

and others present. These forests often occur on north-facing slopes, low slopes, high terraces along streams, and possibly

other situations. Forests in this alliance occur in the low southern Appalachian Mountains, Cumberlands and Southern Ridge and

Valley, Piedmont and Interior Low Plateau, and on protected slopes and ravines in the Ozarks, central Ouachita Mountains, and

Arkansas Valley.

SIMILAR ALLIANCES:

COMMENTS: The relationship between this alliance and I.B.2.N.a.16 Fagus grandifolia - Quercus alba Forest Alliance (A.228)

needs to be clarified. Vegetation from this alliance is known from Ozark and Ouachita national forests RNAs (Roaring Branch

and Dismal Hollow) and occurs on the Shoal Creek District of the Talladega National Forest.

RANGE: This alliance is found in southern Indiana (?), Alabama, Arkansas, Georgia, Kentucky, Louisiana (?), Mississippi, North

Carolina, Oklahoma, South Carolina, Tennessee, Texas (?), Virginia, and West Virginia. Forests in this alliance occur in the low

southern Appalachian Mountains, Cumberlands and Southern Ridge and Valley, Piedmont and Interior Low Plateau, and on

protected slopes and ravines in the Ozarks, central Ouachita Mountains, and Arkansas Valley.

TNC ECOREGIONS: 38:C, 39:C, 43:C, 44:C, 49:C, 50:C, 51:?, 52:C, 57:?, 58:P, 61:P, 62:C

FEDERAL LANDS: NPS (Buffalo, Mammoth Cave, Shiloh); TVA (Land Between the Lakes, Tellico); USFS (Bankhead,

Cherokee?, Conecuh, Daniel Boone, Homochitto, Jefferson?, Ouachita, Ozark, Sumter, Talladega, Tuskegee, Uwharrie)

SYNONYMY: Typic Mesic Piedmont Forest, in part (Allard 1990); Mixed Mesophytic Forest, in part (Foti 1994); Piedmont Mesic

Broad-leaved Deciduous Forest (Ambrose 1990); Mesic Mixed Hardwood Forest, Piedmont subtype (Schafale and Weakley

1990); Appalachian mesophytic forest, in part (Evans 1991); Beech RV? (Pyne 1994); T1B4aI. Fagus grandifolia, TIB4aI1a.

Fagus grandifolia/Magnolia tripetala, T1B4aI1b. Fagus grandifolia - Acer saccharum - Quercus spp. (alba, muehlenbergii,

rubra) (Foti et al. 1994); Beech - Sugar Maple: 60, in part (Eyre 1980); Northern Red Oak: 55, in part (Eyre 1980).

REFERENCES: Allard 1990, Evans 1991, Eyre 1980, Faber-Langendoen et al. 1996, Foti et al. 1994, Golden 1979, Jones

1988a, Jones 1988b, Martin and Smith 1991, Schafale and Weakley 1990, United States Forest Service 1990
Fagus grandifolia - Quercus rubra / Tilia americana var. americana - Ilex opaca Forest (MP 11-98) (CEGL007823 SCS

American Beech - Red Oak / American Basswood - American Holly Forest

American Beech Temporarily Flooded Forest Alliance

CONCEPT: This alliance contains vegetation that occurs along small streams and is dominated by Fagus grandifolia. Other

canopy species include Quercus alba, Quercus rubra var. rubra, Quercus laurifolia, Acer barbatum, Quercus shumardii var.

the Piedmont of North Carolina. Herbaceous species common to forests in this alliance include Carex glaucescens, Carex

this alliance is incomplete, but it is known from at least the Piedmont of North Carolina, Virginia, and South Carolina, the West

Gulf Coastal Plain, southern Indiana, Pennsylvania, and possibly southern Ontario in Canada.

SIMILAR ALLIANCES:

COMMENTS: This alliance needs review. Although these forests experience flooding, perhaps even annually, the vegetation

does not appear to be influenced by these disturbances (i.e. Fagus grandifolia is dominant), so these associations might be

placed better in an upland alliance. Mixed beech-other species forests occur in Texas, but these lack enough Fagus to be

included here. Work is needed to determine alliance placement. The so-called "beechy bottoms" of Kentucky and southern

Illinois, which are high terraces with beech dominant, could be included here. This alliance is present along the Notaway and

Meherrin rivers in Virginia (G. Fleming pers. comm.).

RANGE: This alliance is found in southern Indiana, Georgia, Kentucky (?), Louisiana, Mississippi, North Carolina, South

Carolina, Tennessee (?), Texas (?), Pennsylvania, Virginia, and possibly in Canada in southern Ontario (?).

TNC ECOREGIONS: 41:C, 43:P, 44:C, 45:C, 48:C, 49:C, 50:P, 52:C, 53:C, 56:C, 57:C, 58:P

FEDERAL LANDS: DOD (Fort Gordon); NPS (Kings Mountain); USFS (Angelina?, Kisatchie, Sabine?, Sam Houston?, Uwharrie)

SYNONYMY: American Beech-Southern Magnolia Forest, in part (Diamond 1993); no equivalent (Eyre 1980). Jackson (1979)

listed an American Beech Cover Type in his classification of Indiana plant communities. This appears to be equivalent to this

alliance in the Midwest.

REFERENCES: Diamond 1993, Faber-Langendoen et al. 1996, Fleming pers. comm., Jackson 1979, Martin and Smith 1991

American Beech - (Southern Magnolia) / Witch-hazel / Giant Cane Forest

White Oak - (Water Oak) Forest Alliance

CONCEPT: Mesic and dry-mesic oak - hickory forests of the coastal plain dominated by Quercus alba, and often

codominated by Quercus nigra. These forests lack significant Quercus rubra and Carya ovata, and while Fagus

have Quercus stellata, Prunus serotina var. serotina, Chimaphila maculata, Vaccinium stamineum, and Vaccinium pallidum.

usually is sparse, and species that may occur include Hexastylis arifolia, Elephantopus spp., Chimaphila maculata, Tephrosia

swamp islands, coastal plain slopes, ridgetops, and other dry areas on acidic soils. This alliance occurs throughout the

southeastern coastal plain and likely is distributed sparingly in the Piedmont.

SIMILAR ALLIANCES: Quercus alba-dominated forests in Kentucky and Tennessee are placed in I.B.2.N.a.27 Quercus alba

COMMENTS:

RANGE: This alliance occurs throughout the southeastern Coastal Plain and likely is distributed sparingly in the Piedmont. It is

found in Alabama (?), Arkansas (?), Georgia, Louisiana (?), Mississippi, North Carolina, South Carolina, Tennessee, Texas, and

Virginia.

TNC ECOREGIONS: 31:P, 39:P, 40:?, 41:?, 42:P, 43:C, 52:P, 53:C, 56:C, 57:C, 58:P

FEDERAL LANDS: DOD (Fort Benning); DOE (Savannah River Site); USFS (Angelina, Apalachicola, Bienville, Conecuh,

Croatan, Davy Crockett?, De Soto, Francis Marion, Holly Springs, Homochitto, Kisatchie, Osceola?, Sabine, Sam Houston?, St.

Francis, Sumter?, Tombigbee, Tuskegee)

SYNONYMY: IA8d. Southern Mixed Hardwood Forest, in part (Allard 1990); Coastal Plain Loess Forest, in part (Wieland 1994b); no equivalent (Diamond 1993); TIB4aII. Quercus alba - Mixed Hardwoods (Foti et al. 1994); White Oak: 53, in part (Eyre 1980).

REFERENCES: , Allard 1990, Eyre 1980, Foti et al. 1994, Monk et al. 1990, Schafale and Weakley 1990, Whipple et al. 1981
Quercus alba - Carya alba / Vaccinium elliottii Forest (JEM) (CEGL007224 SCS 390-10)

White Oak - Mockernut Hickory / Mayberry Forest

White Oak - Pignut Hickory - Mockernut Hickory / Red Buckeye Forest

[Basic Mesic Coastal Plain Oak--Hickory Forest]
I.B.2.N.a.27 QUERCUS ALBA - (QUERCUS RUBRA, CARYA SPP.) FOREST ALLIANCE (DJA/DF-L, MOD. SL/DF-L 97-01)

(A.239 MCS )

White Oak - (Red Oak, Hickory species) Forest Alliance

CONCEPT: This alliance, widely distributed in the eastern United States and portions of adjacent Canada, contains drier upland

oak forests. Stands are 15-25 m tall, with a closed, deciduous canopy. The shrub and herbaceous strata are well-developed.

sometimes Quercus velutina (especially on drier sites) or Carya ovata. In the Southeast Region, this alliance covers the

dry-mesic forests of the Piedmont/low Mountain/low Plateau, and mesic oak-hickory forests of interior uplands, which occur

frequently on concave landforms and often on richer soils than forests of the I.B.2.N.a.100 Quercus alba - Quercus velutina -

slopes. Associated species in the Southeast include Carya glabra, Carya ovata, Carya alba, Fraxinus americana, Acer

others. In the Midwest Region, this alliance is found throughout the region on moderately rich, upland sites. Typical associates

include Fraxinus americana, Ulmus americana, Tilia americana, Acer saccharum, Acer rubrum, and more locally, Quercus

macrocarpa and Quercus ellipsoidalis. ^Stands are found on gentle to moderately steep slopes on uplands and on steep

valley sides. The soils are moderately deep to deep and vary from silts to clays and loams. The parent material ranges from

glaciated till to limestone, shale, sandstone and other bedrock types. In the Midwest, many stands are succeeding to types

dominated by Acer saccharum, Tilia americana, Acer rubrum, and other mesic tree associates. This succession may be

delayed by fire and grazing. In the East and Southeast, Liriodendron tulipifera, Fraxinus americana, and other mesic

associates often increase after disturbances, such as clearcutting or windstorms, especially in the absence of fire (Eyre 1980).

SIMILAR ALLIANCES: See the I.B.2.N.a.100 Quercus alba - Quercus velutina - (Quercus coccinea) Forest Alliance (A.1911)

for floristically and structurally similar stands.

COMMENTS:

RANGE: This alliance is found in Alabama, Arkansas, Georgia, Kentucky, North Carolina, Oklahoma, South Carolina,

Tennessee, Connecticut, Massachusetts, Maryland (?), Maine, New Hampshire, New Jersey, New York (?), Pennsylvania (?),

Rhode Island, Virginia, Illinois, Indiana, Iowa, Kansas, Michigan, Minnesota, Missouri, Nebraska, Ohio, and Wisconsin, and in

Canada in Ontario.

TNC ECOREGIONS: 32:P, 35:C, 36:C, 37:C, 38:C, 39:C, 43:C, 44:C, 45:C, 46:C, 47:C, 48:C, 49:C, 50:C, 51:C, 52:C, 53:?,

58:P, 59:P, 61:P, 62:C

FEDERAL LANDS: COE (Dale Hollow?); DOD (Arnold, Fort Benning?); NPS (Chickamauga-Chattanooga, Great Smoky

Mountains; Kennesaw Mountain, Kings Mountain, Russell Cave, Shiloh); TVA (Tellico); USFS (Bankhead, Chattahoochee,

Cherokee?, Daniel Boone, Nantahala, Oconee, Ouachita, Ozark, Pisgah, St. Francis, Sumter, Talladega, Uwharrie)

SYNONYMY: IA6j. Interior Calcareous Oak - Hickory Forest, in part (Allard 1990); Mesic Oak-Hickory Forest, in part (Foti

1994); Submesic broad-leaved deciduous forest, in part (Ambrose 1990); Oak - Chestnut - Hickory Forest, in part (Ambrose

1990); Acidic mesophytic forest, in part (Evans 1991); Calcareous mesophytic forest, in part (Evans 1991); Dry-mesic

Oak--Hickory Forest (Schafale and Weakley 1990); Basic Oak - Hickory Forest, Mafic Substrate Variant, in part (Schafale and

Weakley 1990); Montane Oak--Hickory Forest, in part (Schafale and Weakley 1990); Basic Oak - Hickory Forest (Nelson 1986);

Permesotrophic Forest, in part (Rawinski 1992); Oak--Hickory Forest, in part (Nelson 1986); Quercus rubra - Quercus spp.

(Foti et al. 1994); White Oak - Black Oak - Northern Red Oak: 52, in part (Eyre 1980); White Oak: 53, in part (Eyre 1980). This

alliance is described in part by portions of SAF cover type 52, White Oak-Black Oak-Northern Red Oak, and much of SAF

cover type 53, White Oak (Eyre 1980). Portions of SAF cover type 52 are also included in the Quercus velutina - Quercus

alba Forest Alliance and the Quercus alba - Quercus (coccinea, rubra, velutina) Forest Alliance. Monk et al. (1990) describe a

Quercus alba forest group dominated by Quercus alba, Quercus rubra, and Quercus velutina stretching from Iowa and

Wisconsin to Connecticut and southward into the piedmont in South Carolina, Georgia, Louisiana, and eastern Texas. Their type

includes the Quercus alba - Quercus (coccinea, rubra, velutina) Forest Alliance.

REFERENCES: Allard 1990, Andreu and Tukman 1995, Evans 1991, Eyre 1980, Faber-Langendoen et al. 1996, Foti et al. 1994,

Fountain and Sweeney 1985, Fralish 1988, Fralish et al. 1991, Golden 1979, Hoagland 1997, Jones 1988a, Jones 1988b,

McLeod 1988, Monk et al. 1990, Nelson 1986, Oakley et al. 1995, Oosting 1942, Robertson et al. 1984, Schafale and Weakley

1990, Wharton 1978
Quercus alba - Quercus rubra - Carya (alba, ovata) / Cornus florida Acid Forest (DFL) (CEGL002067 MCS 390-10)

White Oak - Red Oak - (Mockernut Hickory, Shagbark Hickory) / Flowering Dogwood Acid Forest

[Oak - Hickory Dry-mesic Acid Forest]
Quercus alba - Quercus rubra - Quercus muehlenbergii / Cercis canadensis Forest (DFL) (CEGL002070 MCS 390-10)

White Oak - Red Oak - Chinquapin Oak / Redbud Forest

[White Oak - Mixed Oak / Redbud Dry-mesic Alkaline Forest]
Quercus alba / Cornus florida Unglaciated Forest (DFL) (CEGL002066 MCS )

White Oak / Flowering Dogwood Unglaciated Forest

[White Oak / Dogwood Dry-Mesic Forest]
Quercus rubra - Quercus shumardii Forest (BH 10-97) (CEGL004796 SCS 390-10)

Red Oak - Shumard Oak Forest

White Oak - (Southern Red Oak, Post Oak) Forest Alliance

CONCEPT: This alliance contains vegetation that can be described as dry oak and oak - hickory forests. These are usually

dominated by a mixture of Quercus alba and Quercus falcata; Quercus stellata may be dominant or codominant. In

addition, Quercus coccinea, Quercus velutina, Quercus marilandica, Carya alba, Carya glabra, Carya pallida, Carya

carolinae-septentrionalis, Carya ovata, and Fraxinus americana often are present. Common subcanopy and shrub species

include Oxydendrum arboreum, Acer rubrum, Ulmus alata, Juniperus virginiana var. virginiana, Vaccinium arboreum, Cornus

successional forests following logging and/or agricultural cropping. Some examples occur in upland flats and have been called

xerohydric because they occasionally will have standing water in the winter due to a perched water table, but are droughty by

the end of the growing season. Other occurrences are found on well-drained sandy loam or clay loam soils that are often,

although not always, shallow. Karst topography can be found in areas where this alliance occurs. Soils are most often a

well-drained sandy loam, although clay loams are not uncommon. Forests of this alliance may occupy narrow bands of

dry-mesic habitat transitional between lower and midslope mesic communities and xeric ridgetops. This alliance is found in the

Upper East Gulf Coastal Plain, Piedmont, low mountains, and Interior Low Plateau. Distribution in the Atlantic Coastal Plain, East

Gulf Coastal Plain, Upper West Gulf Coastal Plain, and Ouachita Mountains needs assessment. In the Shawnee Hills, Knobs,

coastal plain, and Appalachian Plateau regions of Kentucky, these forests form a common matrix vegetation over acid

sandstone and shales. These Kentucky forests are dominated by Quercus alba with little or no Quercus falcata and occupy

middle to upper slope positions. In the southern Illinois portion of the range, examples occur on south- to west-facing slopes

where increased temperatures favor Quercus falcata over Quercus rubra.

SIMILAR ALLIANCES: These forests are drier than those of the I.B.2.N.a.27 Quercus alba - (Quercus rubra, Carya spp.)

Forest Alliance (A.239) and the I.B.2.N.a.100 Quercus alba - Quercus velutina - (Quercus coccinea) Forest Alliance (A.1911)

and often occur on poorer soils or on south- and west-facing slopes. Related forests, drier than those of this alliance, are placed

in the I.B.2.N.a.31 Quercus falcata Forest Alliance (A.243), of which the range is farther south. This alliance could occur in

the Ozarks and Ouachita Mountains, but associations are yet to be defined. Distribution of vegetation in this alliance is patchy,

occurring on south- to west-facing slopes where Quercus falcata is favored over Quercus rubra. Stands of this alliance are

similar to, and can grade into, stands of other dry-mesic Quercus spp.-dominated alliances such as the I.B.2.N.a.27 Quercus

the Midwest.

COMMENTS:

RANGE: This alliance is found in southern Illinois, Kentucky, Tennessee, Arkansas, Louisiana (?), Oklahoma (?), Texas (?),

Mississippi, Alabama, Georgia, South Carolina, North Carolina, Virginia, Delaware, Maryland, and New Jersey. This alliance is

found in the Upper East Gulf Coastal Plain, Piedmont, low mountains, Interior Low Plateau, and Ouachitas. Distribution in the

Atlantic Coastal Plain, East Gulf Coastal Plain, and Upper West Gulf Coastal Plain needs assessment. In the Shawnee Hills,

Knobs, Coastal Plain, and Appalachian Plateau regions of Kentucky, these forests form a common matrix vegetation over acid

sandstone and shales.

TNC ECOREGIONS: 32:P, 38:?, 39:?, 40:C, 41:P, 42:P, 43:C, 44:C, 50:C, 51:?, 52:C, 53:P, 56:C, 57:P, 58:C, 59:P, 62:C

FEDERAL LANDS: DOD (Arnold, Fort Benning, Fort Gordon); NPS (Big South Fork, Chickamauga-Chattanooga, Kennesaw

Mountain, Shiloh); TVA (Land Between the Lakes?, Tellico); USFS (Bankhead?, Daniel Boone, Holly Springs?, Kisatchie?,

Oconee, Ouachita?, Ozark, Sabine?, St. Francis, Shawnee, Sumter, Talladega, Tombigbee?, Tuskegee?, Uwharrie)

SYNONYMY: IA6i. Interior Upland Dry-Mesic Oak - Hickory Forest, in part (Allard 1990); Acidic sub-xeric forest, in part (Evans

1991); Xerohydric flatwoods, in part (Evans 1991); Dry-Mesic Oak - Hickory Forest, in part (Schafale and Weakley 1990);

Southern Red Oak RV (Pyne 1994); Post Oak-Black Hickory Series, (Diamond 1993); Submesic Oak - Hickory Forest, in part

(Foti 1994); TIB4aIV. Quercus falcata - Quercus spp. (Foti et al. 1994); White Oak - Black Oak - Northern Red Oak: 52, in part

(Eyre 1980). This alliance previously has not been described individually. Braun (1950, p. 154-158) noted the prominence of

Region as well as in the Mississippian Plateau. This alliance was included within SAF cover type 52, White Oak - Black Oak -

Northern Red Oak, as the White Oak - Southern Red Oak variant (Eyre 1980).

REFERENCES: Allard 1990, Andreu and Tukman 1995, Braun 1950, Diamond 1993, Evans 1991, Eyre 1980, Faber-Langendoen et al. 1996, Foti et al. 1994, Fralish et al. 1991, Golden 1979, Oosting 1942, Peet and Christensen 1980, Robertson and Heikens 1994, Schafale and Weakley 1990, Sneddon et al. 1996, Voigt and Mohlenbrock 1964