National Park Service
Northeastern Region
Catalog of Invasive Plants
Invasive Plant Identification Workshop
6-7 August 2002
Delaware Water Gap National Recreation Area
Bushkill Visitor Information Center
Wayne Millington, NPS Project Coordinator
Leslie J. Mehrhoff
Katie R. Boiteau
Stacey A. Leicht
Purpose
The purpose of this project is to identify those plants considered invasive by the National Park Service in the following nine Northeastern states:
Maine, New Hampshire, Vermont, Massachusetts, Rhode Island, Connecticut, New York, New Jersey and Pennsylvania.
This list includes species that are not usually found on most invasive species lists. They are included here because of their potential to invade minimally managed areas within the parks. These species are designated as anthropogenic in the following tables. This list was compiled from existing federal, state and regional lists of invasive species and noxious weeds from the Northeastern United States.
Three tables are included, each containing the same 107 species. These tables are arranged by common name and growth form, scientific name and growth form and scientific name and invasiveness.
In most cases, species names were taken from the Integrated Taxonomic Information System (ITIS) (www.itis.usda.gov). However, there are a few exceptions where, for ease of utility, old names were maintained under species name and currently accepted names were listed under synonyms.
This catalog is a work in progress. If you feel that a species not acknowledged here should be added or have additional information, please let us know.
Please contact:
Leslie J. Mehrhoff
Department of Ecology and Evolutionary Biology
University of Connecticut
Box U-43
75 North Eagleville Road
Storrs, CT 06269-3043
(860) 486-5708
FAX (860) 486-6364
vasculum@uconnvm.uconn.edu
or
Katie Boiteau
krboiteau@hotmail.com
SPECIES NAME
Acer platanoides L.
COMMON NAME
Norway maple
FAMILY
Aceraceae
NPS CODE
ACEPLA
NPS NORTHEASTERN STATUS
Invasive
THREAT
Has dense foliage that lasts longer than most native species. Prohibits light from reaching woodland floor, and produces many seedlings.
INVASIVE ECOLOGY
Grows rapidly when young. Well adapted to extreme soils. Creates very dense shade and seeds freely, causing many weed seedling trees. Very dense foliage. Will withstand sand, clay, acidic to calcareous soils, can withstand hot, dry conditions, tolerates polluted atmosphere. Seedlings are shade tolerant and thrive in hedge rows and roadside thickets. Seeds are much heavier than sugar maple seeds, giving them larger reserves for germination. Mature Acer platanoides facilitate the growth of their seedlings underneath their canopy. Possibly secrete an allelopathic chemical from their roots.
HABITAT
Common street and ornamental tree, found in woodlots and urban-fringe forests. Spreads to successional forests.
GEOGRAPHIC ORIGIN
Eurasia
HISTORY OF INTRODUCTION
Introduced to the United States in 1756; later escaped from cultivation.
NORTHEASTERN US DISTRIBUTION
Entire Northeast
NORTHEASTERN NATIONAL PARKS
ACAD, DEWA, GATE
REPRODUCTION
Deciduous tree, flowers in April. Insect pollinated. Vigorous reproduction from seed.
FRUIT AND SEED PRODUCTION
Fruits mature in September to October. Seeds are wind dispersed.
CONTROL
Girdling or herbicide application to cut stems.
SPECIES NAME
Acer pseudoplatanus L.
COMMON NAME
Sycamore maple
FAMILY
Aceraceae
NPS CODE
ACEPSE
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Seeds germinate easily producing many individuals, especially in sandy soils near the coast.
INVASIVE ECOLOGY
Easily established and very adaptable. Tolerant of salt, high pH, calcareous soil and exposed areas.
GEOGRAPHIC ORIGIN
Europe and Asia
HISTORY OF INTRODUCTION
Introduced to the United States as an ornamental.
NORTHEASTERN US DISTRIBUTION
Maine, Massachusetts, Connecticut, Rhode Island, New York, Pennsylvania, New Jersey, Maryland
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Deciduous tree, flowers in May. Reproduces from seed.
FRUIT AND SEED PRODUCTION
Fruits mature in August or September
CONTROL
SPECIES NAME
Acer palmatum Thunb.
COMMON NAME
Japanese maple
FAMILY
Aceraceae
NPS CODE
ACEPAL
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Produces many seedlings in some areas.
INVASIVE ECOLOGY
Spreads rapidly by seed.
GEOGRAPHIC ORIGIN
Japan, China, Korea
HISTORY OF INTRODUCTION
Introduced from Japan to England in 1840, then to the U.S. in 1862 by Dr. George Hall to a nursery in Flushing, N.Y.
NORTHEASTERN US DISTRIBUTION
New York and Pennsylvania
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Deciduous tree, flowers in May and June. Reproduces from seed.
FRUIT AND SEED PRODUCTION
Fruits ripen in September to October.
CONTROL
SPECIES NAME
Aegopodium podagraria L.
COMMON NAME
Goutweed
FAMILY
Apiaceae
NPS CODE
AEGPOD
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Spreads easily by fragmenting rhizomes and seeds.
INVASIVE ECOLOGY
Grows well in flood plan forests, waste places and roadsides. Weedy and difficult to control.
GEOGRAPHIC ORIGIN
Eurasia
HISTORY OF INTRODUCTION
Cultivated in gardens and often escaped in New England.
NORTHEASTERN US DISTRIBUTION
Entire Northeast
NORTHEASTERN NATIONAL PARKS
DEWA, WEFA
REPRODUCTION
Herbaceous perennial, flowers in June. Reproduces vegetatively by rhizomes and from seed.
FRUIT AND SEED PRODUCTION
Fruits mature in July.
CONTROL
SPECIES NAME
Ailanthus altissima (P. Mill.) Swingle
COMMON NAME
Tree of Heaven
FAMILY
Simaroubaceae
SYNONYMS
Ailanthus glandulosa Desf.
NPS CODE
AILALT
NPS NORTHEASTERN STATUS
Invasive
THREAT
Produces many seedlings. Also spreads vegetatively.
INVASIVE ECOLOGY
Rapid growth, does well in poor soil and with atmospheric pollution. Tolerant of salt, poor soil, heat and drought. Can sucker and produce stump sprouts, and produces large quantities of seeds. Fruits persist through winter.
HABITAT
Woodlots, along railways and highways and other disturbed areas.
GEOGRAPHIC ORIGIN
China
HISTORY OF INTRODUCTION
Introduced to the U.S. in Pennsylvania by a gardener. By 1840, was being sold from nurseries, and has since been uses extensively for plantings in cities.
NORTHEASTERN US DISTRIBUTION
Found in all Northeastern states
NORTHEASTERN NATIONAL PARKS
MIMA, GATE, ROVA, SAHI, DEWA
REPRODUCTION
Deciduous tree, flowers in late spring. Reproduces from seed.
FRUIT AND SEED PRODUCTION
Fruits appear in September to October. Seeds are wind and water dispersed.
CONTROL
Hand-pulling, especially by the Bradley Method, cutting, digging, girdling, prescribed burning, herbicide and grazing until seedbank is eliminated.
SPECIES NAME
Akebia quinata (Houtt.) Dcne.
COMMON NAME
Fiveleaf akebia, Chocolate vine
FAMILY
Lardizabalaceae
NPS CODE
AKEQUI
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Rampant grower with long stems than run over the ground or climb.
INVASIVE ECOLOGY
Can grow upright along a structure or along the ground as a groundcover. Tolerant of many soil types, easily grown. Able to grow rampantly. Appears to produces fruits infrequently.
GEOGRAPHIC ORIGIN
China, Korea, Japan
HISTORY OF INTRODUCTION
Introduced to the United States in 1845.
NORTHEASTERN US DISTRIBUTION
Entire Northeast except Maine, New Hampshire and Vermont.
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Twining woody vine, flowers in late March to early April. Reproduces vegetatively.
FRUIT AND SEED PRODUCTION
Fruits ripen in September to October, each hold numerous seeds. Dispersal mechanisms are unknown, but probably by animals.
CONTROL
Cutting, digging, and herbicides.
SPECIES NAME
Albizia julibrissin Durazz.
COMMON NAME
Silk tree, Mimosa
FAMILY
Fabaceae
NPS CODE
ALBJUL
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Produces many seedlings.
INVASIVE ECOLOGY
Will withstand drought, high pH, soil salinity, and excessive wind. Produce large seed crops, can resprout when damaged. Strongly competitive to native tress and shrubs in open areas or edges. Can form dense stands.
GEOGRAPHIC ORIGIN
Iran to central China.
HISTORY OF INTRODUCTION
Introduced to the United States in 1745.
NORTHEASTERN US DISTRIBUTION
New York, Pennsylvania, New Jersey, Maryland, West Virginia, Virginia. Cultivated in Massachusetts and Connecticut.
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Deciduous tree, flowers in late April through early July. Reproduces from seed or by root cuttings.
FRUIT AND SEED PRODUCTION
Fruits ripen in August to October and persist through the winter often into the spring. Seeds dispersed around parent plant and by water.
CONTROL
Cutting at ground level, herbicides, girdling, hand pulling of young seedlings
SPECIES NAME
Alliaria petiolata (Bieb.) Cavara & Grande
COMMON NAME
Garlic mustard
FAMILY
Brassicaceae
SYNONYMS
Alliaria alliaria (L.) Britt., Alliaria officinalis Andrz. ex Bieb.
NPS CODE
ALLPET
NPS NORTHEASTERN STATUS
Invasive
THREAT
Produces large numbers of individuals that can form dense monotypic stands even in closed canopy forests. Persists in seed banks.
INVASIVE ECOLOGY
Shade tolerant and can dominate a forest understory. Tolerates a wide variety of soils and moisture levels. Dense stands Deprive native herbaceous species of light, moisture and space. High seed production and strong seed dormancy
HABITAT
Common in forest edges, roadsides, stream banks, and gardens in shaded and semi-shaded habitats.
GEOGRAPHIC ORIGIN
Europe
HISTORY OF INTRODUCTION
First collected in Long Island in 1868. Likely brought to the United States as a food crop and for medicinal purposes.
NORTHEASTERN US DISTRIBUTION
Entire Northeast
NORTHEASTERN NATIONAL PARKS
ACAD, ALPO, DEWA, ROVA
REPRODUCTION
Biennial herb, flowers in the spring. Reproduces from seed.
FRUIT AND SEED PRODUCTION
Fruits produced by May, with 10-20 seeds per pod by early summer. Seeds are mechanically dispersed, and dispersed by humans and animals.
CONTROL
Prevention of initial establishment, cutting the flowering stems at ground level annually, and biocontrol.
SPECIES NAME
Alnus glutinosa (L.) Gaertn.
COMMON NAME
European black alder
FAMILY
Betulaceae
SYNONYMS
Alnus alnus (L.) Britt., Betula glutinosa L.
NPS CODE
ALNGLU
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Can form dense stands of many individuals.
INVASIVE ECOLOGY
Fast growth rate in youth, tolerant of acid or slightly alkaline soils, tolerant of frost, poor soil and waterlogging.
GEOGRAPHIC ORIGIN
Europe and western Asia
HISTORY OF INTRODUCTION
Cultivated as an ornamental, and planted to control erosion and improve the soil on recently cleared or unstable substrates.
NORTHEASTERN US DISTRIBUTION
Connecticut, Massachusetts, New Jersey, New York, Pennsylvania, Rhode Island, Vermont
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Deciduous tree. Reproduces from seeds and spreads vegetatively.
FRUIT AND SEED PRODUCTION
Fruits mature in October and November
CONTROL
Girdling or cutting stems and applying herbicide
SPECIES NAME
Ampelopsis brevipedunculata (Maxim.) Trautv.
COMMON NAME
Porcelain berry
FAMILY
Vitaceae
NPS CODE
AMPBRE
NPS NORTHEASTERN STATUS
Invasive
THREAT
Vines grow rampantly up into the tree canopy or over other vegetation.
INVASIVE ECOLOGY
Can grow over and smother native vegetation. Rapid growth, with a high germination rate.
HABITAT
Woodland edges, riparian areas, and open fields, especially areas with repeated disturbance.
GEOGRAPHIC ORIGIN
Japan, China, Korea, and the Russian Far East
HISTORY OF INTRODUCTION
Introduced into cultivation in the U.S. for use as a bedding and screening plant.
NORTHEASTERN US DISTRIBUTION
Maine, New Hampshire, Vermont, Massachusetts, Connecticut and Rhode Island
NORTHEASTERN NATIONAL PARKS
MIMA
REPRODUCTION
Deciduous, perennial vine, flowers in summer. Reproduces from seed and cuttings.
FRUIT AND SEED PRODUCTION
Fruits ripen in September and contain 2-4 seeds each. Seeds are bird and water dispersed.
CONTROL
Hand pulling or pruning before fruiting and cutting aboveground vines. Herbicides can be applied to leaves or to basal bark.
SPECIES NAME
Anthriscus sylvestris (L.) Hoffmann
COMMON NAME
Wild chervil
FAMILY
Apiaceae
NPS CODE
ANTSYL
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Produces many seeds that can form dense stands of many individuals.
INVASIVE ECOLOGY
Self-seeds, has long taproot, grows well in woodlands, forest, edges, waste places and roadsides.
GEOGRAPHIC ORIGIN
Europe
HISTORY OF INTRODUCTION
NORTHEASTERN US DISTRIBUTION
Entire Northeast
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Biennial or short-lived perennial forb, flowers in May through July. Pollinated by bees.
FRUIT AND SEED PRODUCTION
Seeds are mature from June through July.
CONTROL
SPECIES NAME
Artemisia stelleriana Bess.
COMMON NAME
Dusty miller
FAMILY
Asteraceae
NPS CODE
ARTSTE
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Vegetative growth can produce large stands in sandy soil along coastal dunes and beaches.
INVASIVE ECOLOGY
Fast growth rate including rhizomes; can tolerate poor soils including sand and rocks
HABITAT
Sandy beaches, coastal sand dunes.
GEOGRAPHIC ORIGIN
Asia
HISTORY OF INTRODUCTION
Introduced to the United States as garden material and escaped from cultivation.
NORTHEASTERN US DISTRIBUTION
Entire Northeast
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Perennial, flowers in summer. Spreads vegetatively by segments rooting in soil.
FRUIT AND SEED PRODUCTION
CONTROL
SPECIES NAME
Artemisia vulgaris L.
COMMON NAME
Mugwort
FAMILY
Asteraceae
NPS CODE
ARTVUL
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Can form dense stands by vegetative growth.
INVASIVE ECOLOGY
Able to grow in a wide variety of soils, prolific seeder and seeds may persist in the soil for many years. Rapid colonizer, crowds out existing vegetation.
HABITAT
Fields, roadsides, disturbed urban areas, river banks and waste places.
GEOGRAPHIC ORIGIN
Europe and Asia
HISTORY OF INTRODUCTION
NORTHEASTERN US DISTRIBUTION
Entire Northeast
NORTHEASTERN NATIONAL PARKS
ACAD, DEWA
REPRODUCTION
Perennial herb, flowers in late summer and early fall. Reproduces from seed and vegetatively by rhizomes.
FRUIT AND SEED PRODUCTION
Viable seeds rarely produced? Seeds dispersed by wind and humans.
CONTROL
Herbicide
SPECIES NAME
Arthraxon hispidus (Thunb.) Makino
COMMON NAME
Hairy joint-grass
FAMILY
Poaceae
SYNONYMS
Phalaris hispida Thunb.
NPS CODE
ARTHIS
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Produces dense monotypic stands annually.
INVASIVE ECOLOGY
Can survive and germinate after extended flooding. Forms monotypic stands and crowds out native herbaceous vegetation. Able to grow in shade. Rapidly spreads along stream and pond shores and through ditches and wet areas.
GEOGRAPHIC ORIGIN
Asia
HISTORY OF INTRODUCTION
May have been introduced with packing material at the Philadelphia Exposition in 1876.
NORTHEASTERN US DISTRIBUTION
South of and including Connecticut.
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Annual grass. Reproduces from seed.
FRUIT AND SEED PRODUCTION
Produces abundant seed.
CONTROL
Hand pulling or mowing before seed production for seven years, herbicide
SPECIES NAME
Arundo donax L.
COMMON NAME
Giant reed
FAMILY
Poaceae
SYNONYMS
Arundo versicolor P. Mill.
NPS CODE
ARUDON
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Forms dense stands by vegetative growth.
INVASIVE ECOLOGY
Salinity tolerant, creates sand dunes, tolerant of all soils. Rootstocks form compact masses, has deeply penetrating fibrous roots. Chokes riversides and stream channels. Crowds out native plants, interferes with flood control. Increases fire potential, decreases wildlife habitat, and can outcompete native vegetation. Rapid growth rate.
GEOGRAPHIC ORIGIN
Mediterranean
HISTORY OF INTRODUCTION
Introduced intentionally to Los Angeles, CA in the early 1800s as an ornamental and for erosion control. Used for making pipe organs, baskets, fishing rods, medicines, and in fodder.
NORTHEASTERN US DISTRIBUTION
West Virginia and Virginia
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Perennial grass, flowers in August and September. Fragments root after floating downstream. Stem fragments can take root. Vegetative and rhizome reproduction. Wind and water dispersed.
FRUIT AND SEED PRODUCTION
Little known
CONTROL
Herbicide, prescribed burning
SPECIES NAME
Berberis thunbergii DC.
COMMON NAME
Japanese barberry
FAMILY
Berberidaceae
NPS CODE
BERTHU
NPS NORTHEASTERN STATUS
Invasive
THREAT
Dense, near monotypic, stands exclude other species and many change the soil chemistry.
INVASIVE ECOLOGY
Form dense, continuous stands. Exceptionally tolerant and adaptable, withstands drought and heat, invades undisturbed as well as disturbed areas. Hybridizes with Berberis vulgaris to form B. X ottawensis.
HABITAT
Roadsides, trails, open fields, gardens and forest edges.
GEOGRAPHIC ORIGIN
Japan
HISTORY OF INTRODUCTION
Introduced to the United States as an ornamental in 1875 as seeds sent from Russia to the Arnold Arboretum in Boston. Planted in the New York Botanic Garden in 1896. Later planted as a substitute for the black stem grain rust infested Berberis vulgaris in seaside gardens. Recognized as a garden escape before 1910.
NORTHEASTERN US DISTRIBUTION
All of Northeast except the Adirondaks, northern Maine and northern Vermont.
NORTHEASTERN NATIONAL PARKS
ACAD, ALPO, DEWA, MABI, MORR, ROVA, WEFA
REPRODUCTION
Deciduous shrub, flowers from April to May. Reproduces by cuttings and from seed. Reproduces when branches come in contact with the soil.
FRUIT AND SEED PRODUCTION
Numerous berries mature from July to October and persist on the stems until spring. Dispersed by turkey, grouse and small mammals.
CONTROL
Pulling by hand, mowing or cutting and herbicide.
SPECIES NAME
Berberis vulgaris L.
COMMON NAME
Barberry
FAMILY
Berberidaceae
NPS CODE
BERVUL
NPS NORTHEASTERN STATUS
Potentially invasive
INVASIVE ECOLOGY
Forms dense thickets, produces abundant seeds, able to grow in shaded habitats. Hybridizes with Berberis thunbergii to create B. X ottawensis.
HABITAT
Pastures, open woodlands and waste places.
GEOGRAPHIC ORIGIN
Europe
HISTORY OF INTRODUCTION
Introduced to the United States in the 17th century, it was planted by settlers for its fruits. Once quite common in the Northeast.
NORTHEASTERN US DISTRIBUTION
All of Northeast
NORTHEASTERN NATIONAL PARKS
ACAD
REPRODUCTION
Deciduous shrub, flowers May to June. Reproduces by seed and vegetatively, and new plants can be produced when branches come in contact with the soil.
FRUIT AND SEED PRODUCTION
Fruits from May to June. Fruits contain 1-3 seeds each. Seeds are dispersed by birds and small mammals.
CONTROL
Pulling by hand, mowing, cutting and herbicide. Many individuals were eradicated by the CCC in the 1920s; in some places many can still be found.
SPECIES NAME
Butomus umbellatus L.
COMMON NAME
Flowering rush
FAMILY
Butomaceae
NPS CODE
BUTUMB
NPS NORTHEASTERN STATUS
Potentially invasive
THREAT
Can invade shore communities with lots of individuals. May form dense monotypic stands under some conditions.
INVASIVE ECOLOGY
Grows well in floodplain forests, aquatic, rivers or streams, lake or pond margins. Now used in water garden habitats. Can displace native riparian vegetation, and has a wide hardiness zone.
GEOGRAPHIC ORIGIN
East Asia
HISTORY OF INTRODUCTION
First observed in 1897 in Quebec. In 1929 collected in two Vermont counties. By 1950, well established along St. Lawrence River. First collected in Connecticut in 1943.
NORTHEASTERN US DISTRIBUTION
Maine, Vermont, Connecticut, New York, Pennsylvania
NORTHEASTERN NATIONAL PARKS
REPRODUCTION
Aquatic plant. Reproduces by seed or vegetatively.
FRUIT AND SEED PRODUCTION
Fruits split at maturity releasing seeds. Seeds are water dispersed.
CONTROL
Cutting below the water surface and hand digging
SPECIES NAME
Cabomba caroliniana Gray
COMMON NAME
Fanwort
FAMILY
Cabombaceae
NPS CODE
CABCAR
NPS NORTHEASTERN STATUS
Invasive
THREAT
Forms dense monotypic stands that prohibit sunlight penetration.
INVASIVE ECOLOGY
Grows well in aquatic, river or stream, lake or pond and water garden habitats. Can form extremely dense stands and clog drainage systems. Able to root from vegetative parts.
HABITAT
Ponds, lakes ditches and quiet streams.
GEOGRAPHIC ORIGIN
Southern United States
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